Gradient function of the normal model
1  | grad_normal(obs, a, Va, Ve, Ut)
 | 
obs | 
 numeric, observed fitnesses  | 
a | 
 numeric, mean effect-size of mutations  | 
Va | 
 numeric variance of dfe  | 
Ve | 
 numeric variance of experimental system  | 
Ut | 
 numeric mutation rate  | 
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