\noindent{} 1. Groningen Institute for Evolutionary Life Sciences, University of Groningen, Groningen 9747 AG, The Netherlands.
\noindent{} $\ast$ Corresponding authors; e-mail: p.r.gupte@rug.nl or f.j.weissing@rug.nl
\noindent{} $\alpha$ Both authors contributed equally to this study.
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