knitr::opts_chunk$set(echo = TRUE)
Variability of the responses:
1. Random drift due to the restricted number of parents;
2. Sampling error in estimating hte generation mean;
3. Variation of selection differentials;
4. Environmental factors.
$$
\sigma_R^2 = \sigma_d^2 + \sigma_e^2
= V_p(th^2/N_e + 1/M)
$$
In the equation, M denotes the of individuals measured.
Selection experiments often yield no response over the first one or two generations, or even longer, but give a clear response later. The reason is usually that the numbers have been too small and random drift in the 'wrong' direction has nullified the response.
c=5 i=1.271 h2=0.3 2*c^2/(i*h2)
Random Drift
to do: simulate the response by considering drift.
Inbreeding Depression
It is that if the character selected is a component of natural fitness, asymmetry should be expected, with selection towards increased fitness giving a slower response than selection towards decreased fitness.
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