The density dependence in this model is roughly a Beverton-Holt model that includes both interspecific and intraspecific competition. Each individual has a random chance of survival based on a variety of conditions. Competition coefficients depend on interactions between each species and the temperature at the location at the time. These can be thought of as temperature-varying Lotka-Volterra competition coefficients. Probability of survival depends on competition coefficients, number of individuals of each different species at that location, and the quality of the habitat at that location.
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