Nothing

```
get.ave.density <- function(density.surface, coords, gaps, x.space, y.space){
#Calculates the average density across the study region
grid.cell.weighting <- function(centre, coords, gaps, x.space, y.space){
#Get 4 corner points
corners <- data.frame(x = (centre[["x"]]+c(x.space, x.space, -x.space, -x.space)), y = (centre[["y"]]+c(y.space, -y.space, y.space, -y.space)))
#Check which have one or more that falls outside the strata
in.strata <- in.polygons(poly.list = coords, pts = corners, boundary = TRUE)
in.gaps <- in.polygons(poly.list = gaps, pts = corners, boundary = TRUE)
if(length(which(in.strata)) == 4 & length(which(in.gaps)) == 0){
return(1)
}else{
#populate with points
x <- seq(centre[["x"]] - x.space, centre[["x"]] + x.space, length = 10)
y <- seq(centre[["y"]] - x.space, centre[["y"]] + x.space, length = 10)
prop.points <- expand.grid(x = x, y = y)
#count what proportion are inside
in.strata <- in.polygons(poly.list = coords, pts = prop.points, boundary = TRUE)
in.gaps <- in.polygons(poly.list = gaps, pts = prop.points, boundary = TRUE)
to.count <- ifelse(in.gaps, FALSE, in.strata)
proportion.in.strata <- sum(to.count)/length(to.count)
return(proportion.in.strata)
}
}
x.space <- x.space/2
y.space <- y.space/2
density.surface <- as.matrix(density.surface)
#takes those which are not entirely within the strata
weighting <- apply(density.surface, 1, FUN = grid.cell.weighting, coords = coords, gaps = gaps, x.space = x.space, y.space = y.space)
#calculate average density
ave.density <- sum(weighting*density.surface[,"density"])/sum(weighting)
return(ave.density)
}
```

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