#' Sets up parameters for a community-type model
#'
#' This functions creates a \code{\linkS4class{therMizerParams}} object so that
#' community-type models can be easily set up and run. A community model has
#' several features that distinguish it from the food-web type models. Only one
#' 'species' is resolved, i.e. one 'species' is used to represent the whole
#' community. The resource spectrum only extends to the start of the community
#' spectrum. Recruitment to the smallest size in the community spectrum is
#' constant and set by the user. As recruitment is constant, the proportion of
#' energy invested in reproduction (the slot \code{psi} of the returned
#' \code{therMizerParams} object) is set to 0. Standard metabolism has been turned
#' off (the parameter \code{ks} is set to 0). Consequently, the growth rate is
#' now determined solely by the assimilated food (see the package vignette for
#' more details).
#'
#' The function has many arguments, all of which have default values. The main
#' arguments that the users should be concerned with are \code{z0},
#' \code{recruitment}, \code{alpha} and \code{f0} as these determine the average
#' growth rate of the community.
#'
#' Fishing selectivity is modelled as a knife-edge function with one parameter,
#' \code{knife_edge_size}, which determines the size at which species are
#' selected.
#'
#' The resulting \code{therMizerParams} object can be projected forward using
#' \code{project_therMizer()} like any other \code{therMizerParams} object. When projecting
#' the community model it may be necessary to reduce \code{dt} to 0.1 to avoid
#' any instabilities with the solver. You can check this by plotting the biomass
#' or abundance through time after the projection.
#'
#' @param z0 The background mortality of the community. Default value is 0.1.
#' @param alpha The assimilation efficiency of the community. Default value 0.2
#' @param f0 The average feeding level of individuals who feed mainly on the
#' resource. This value is used to calculate the search rate parameter
#' \code{gamma} (see the package vignette). Default value is 0.7.
#' @param h The maximum food intake rate. Default value is 10.
#' @param beta The preferred predator prey mass ratio. Default value is 100.
#' @param sigma The width of the prey preference. Default value is 2.0.
#' @param q The search volume exponent. Default value is 0.8.
#' @param n The scaling of the intake. Default value is 2/3.
#' @param kappa The carrying capacity of the background spectrum. Default value
#' is 1000.
#' @param lambda The exponent of the background spectrum. Default value is 2 + q
#' - n.
#' @param r_pp Growth rate of the primary productivity. Default value is 10.
#' @param gamma Volumetric search rate. Estimated using \code{h}, \code{f0} and
#' \code{kappa} if not supplied.
#' @param recruitment The constant recruitment in the smallest size class of the
#' community spectrum. This should be set so that the community spectrum
#' continues the background spectrum. Default value = \code{kappa} *
#' \code{min_w}^-\code{lambda}.
#' @param rec_mult Additional multiplier for the constant recruitment. Default
#' value is 1.
#' @param knife_edge_size The size at the edge of the knife-selectivity
#' function. Default value is 1000.
#' @param knife_is_min Is the knife-edge selectivity function selecting above
#' (TRUE) or below (FALSE) the edge. Default is TRUE.
#' @param max_w The maximum size of the community. The \code{w_inf} of the
#' species used to represent the community is set to 0.9 * this value. The
#' default value is 1e6.
#' @param min_w The minimum size of the community. Default value is 1e-3.
#' @param ... Other arguments to pass to the \code{therMizerParams} constructor.
#' @export
#' @return An object of type \code{\linkS4class{therMizerParams}}
#' @references K. H. Andersen,J. E. Beyer and P. Lundberg, 2009, Trophic and
#' individual efficiencies of size-structured communities, Proceedings of the
#' Royal Society, 276, 109-114
#' @examples
#' \dontrun{
#' params <- set_community_model(f0=0.7, z0=0.2, recruitment=3e7)
#' sim <- project_therMizer(params, effort = 0, t_max = 100, dt=0.1)
#' plotBiomass(sim)
#' plotSpectra(sim)
#' }
set_community_model <- function(max_w = 1e6,
min_w = 1e-3,
z0 = 0.1,
alpha = 0.2,
h = 10,
beta = 100,
sigma = 2.0,
q = 0.8,
n = 2/3,
kappa = 1000,
lambda = 2+q-n,
f0 = 0.7,
r_pp = 10,
gamma = NA,
knife_edge_size = 1000,
knife_is_min = TRUE,
recruitment = kappa * min_w^-lambda,
rec_mult = 1,
...
){
w_inf <- max_w * 0.9
w_pp_cutoff <- min_w
ks <- 0 # Turn off standard metabolism
p <- n # But not used as ks = 0
# Estimate gamma if not supplied
if (is.na(gamma)){
gamma <- (f0 * h * beta^(2-lambda)) / ((1-f0)*sqrt(2*pi)*kappa*sigma)
}
# Make the species data.frame
com_params_df <- data.frame(
species = "Community",
w_inf = w_inf,
w_mat = 1e12, # Has no affect as psi set to 0 but we set it to something
# to help the constructor
h = h, # max food intake
gamma = gamma,# vol. search rate,
ks = ks,# standard metabolism coefficient,
beta = beta,
sigma = sigma,
z0 = z0, # background mortality
alpha = alpha,
erepro = 1, # not used
sel_func = "knife_edge",
knife_edge_size = knife_edge_size,
knife_is_min = knife_is_min,
constant_recruitment = recruitment * rec_mult # to be used in the SRR
)
# Set the recruitment function for constant recruitment
constant_recruitment <- function(rdi, species_params){
return(species_params$constant_recruitment)
}
com_params <- therMizerParams(com_params_df, p=p, n=n,q=q, lambda = lambda,
kappa = kappa, min_w = min_w, max_w = max_w,
w_pp_cutoff = w_pp_cutoff, r_pp = r_pp, ...)
com_params@srr <- constant_recruitment
com_params@psi[] <- 0 # Need to force to be 0. Can try setting w_mat but
# due to slope still not 0
# Set w_mat to NA for clarity - it is not actually being used
com_params@species_params$w_mat[] <- NA
return(com_params)
}
#' Sets up parameters for a trait-based model
#'
#' This functions creates a \code{therMizerParams} object so that trait-based-type
#' models can be easily set up and run. The trait-based size spectrum model can
#' be derived as a simplification of the general size-based model used in
#' \code{therMizer}. All the species-specific parameters are the same, except for
#' the asymptotic size, which is considered the most important trait
#' characterizing a species. Other parameters are related to the asymptotic
#' size. For example, the size at maturity is given by w_inf * eta, where eta is
#' the same for all species. For the trait-based model the number of species is
#' not important. For applications of the trait-based model see Andersen &
#' Pedersen (2010). See the \code{mizer} vignette for more details and examples
#' of the trait-based model.
#'
#' The function has many arguments, all of which have default values. Of
#' particular interest to the user are the number of species in the model and
#' the minimum and maximum asymptotic sizes. The asymptotic sizes of the species
#' are spread evenly on a logarithmic scale within this range.
#'
#' The stock recruitment relationship is the default Beverton-Holt style. The
#' maximum recruitment is calculated using equilibrium theory (see Andersen &
#' Pederson, 2010) and a multiplier, \code{k0}. Users should adjust \code{k0} to
#' get the spectra they want.
#'
#' The factor for the search volume, \code{gamma}, is calculated using the
#' expected feeding level, \code{f0}.
#'
#' Fishing selectivity is modelled as a knife-edge function with one parameter,
#' \code{knife_edge_size}, which is the size at which species are selected. Each
#' species can either be fished by the same gear (\code{knife_edge_size} has a
#' length of 1) or by a different gear (the length of \code{knife_edge_size} has
#' the same length as the number of species and the order of selectivity size is
#' that of the asymptotic size).
#'
#' The resulting \code{therMizerParams} object can be projected forward using
#' \code{project_therMizer()} like any other \code{therMizerParams} object. When projecting
#' the community model it may be necessary to reduce \code{dt} to 0.1 to avoid
#' any instabilities with the solver. You can check this by plotting the biomass
#' or abundance through time after the projection.
#' @param no_sp The number of species in the model. The default value is 10. The
#' more species, the longer takes to run.
#' @param min_w_inf The asymptotic size of the smallest species in the
#' community.
#' @param max_w_inf The asymptotic size of the largest species in the community.
#' @param no_w The number of size bins in the community spectrum.
#' @param min_w The smallest size of the community spectrum.
#' @param max_w The largest size of the community spectrum. Default value is the
#' largest w_inf in the community x 1.1.
#' @param min_w_pp The smallest size of the background spectrum.
#' @param no_w_pp The number of the extra size bins in the background spectrum
#' (i.e. the difference between the number of sizes bins in the community
#' spectrum and the full spectrum).
#' @param w_pp_cutoff The cut off size of the background spectrum. Default value
#' is 1.
#' @param k0 Multiplier for the maximum recruitment. Default value is 50.
#' @param n Scaling of the intake. Default value is 2/3.
#' @param p Scaling of the standard metabolism. Default value is 0.75.
#' @param q Exponent of the search volume. Default value is 0.9.
#' @param eta Factor to calculate \code{w_mat} from asymptotic size.
#' @param r_pp Growth rate of the primary productivity. Default value is 4.
#' @param kappa Carrying capacity of the resource spectrum. Default value is
#' 0.005.
#' @param lambda Exponent of the resource spectrum. Default value is (2+q-n).
#' @param alpha The assimilation efficiency of the community. The default value
#' is 0.6
#' @param ks Standard metabolism coefficient. Default value is 4.
#' @param z0pre The coefficient of the background mortality of the community. z0
#' = z0pre * w_inf ^ (n-1). The default value is 0.6.
#' @param h Maximum food intake rate. Default value is 30.
#' @param beta Preferred predator prey mass ratio. Default value is 100.
#' @param sigma Width of prey size preference. Default value is 1.3.
#' @param f0 Expected average feeding level. Used to set \code{gamma}, the
#' factor for the search volume. The default value is 0.5.
#' @param gamma Volumetric search rate. Estimated using \code{h}, \code{f0} and
#' \code{kappa} if not supplied.
#' @param knife_edge_size The minimum size at which the gear or gears select
#' species. Must be of length 1 or no_sp.
#' @param gear_names The names of the fishing gears. A character vector, the
#' same length as the number of species. Default is 1 - no_sp.
#' @param ... Other arguments to pass to the \code{therMizerParams} constructor.
#' @export
#' @return An object of type \code{therMizerParams}
#' @seealso \linkS4class{therMizerParams}
#' @references K. H. Andersen and M. Pedersen, 2010, Damped trophic cascades
#' driven by fishing in model marine ecosystems. Proceedings of the Royal
#' Society V, Biological Sciences, 1682, 795-802.
#' @examples
#' \dontrun{
#' trait_params <- set_trait_model(no_sp = 15)
#' init_pop <- get_initial_n(trait_params, n0_mult = 0.001)
#' sim <- project_therMizer(trait_params, effort = 0, t_max = 50, dt=0.2,
#' initial_n = init_pop, t_save = 1)
#' plot(sim)
#' ## Set up industrial fishery that only fishes on species with w_inf <= 500 g
#' ## And where the selectivity of the industrial fishery = w_inf * 0.05
#' no_sp <- 10
#' min_w_inf <- 10
#' max_w_inf <- 1e5
#' w_inf <- 10^seq(from=log10(min_w_inf), to = log10(max_w_inf), length=no_sp)
#' knife_edges <- w_inf * 0.05
#' industrial_gears <- w_inf <= 500
#' other_gears <- w_inf > 500
#' gear_names <- rep("Industrial", no_sp)
#' gear_names[other_gears] <- "Other"
#' params_gear <- set_trait_model(no_sp = no_sp, min_w_inf = min_w_inf,
#' max_w_inf = max_w_inf, knife_edge_size = knife_edges, gear_names = gear_names)
#' ## Only turn on Industrial fishery. Set effort of the Other gear to 0
#' sim <- project_therMizer(params_gear, t_max = 20, effort = c(Industrial = 1, Other = 0))
#' }
set_trait_model <- function(no_sp = 10,
min_w_inf = 10,
max_w_inf = 1e5,
no_w = 100,
min_w = 0.001,
max_w = max_w_inf * 1.1,
min_w_pp = 1e-10,
no_w_pp = round(no_w)*0.3,
w_pp_cutoff = 1,
k0 = 50, # recruitment adjustment parameter
n = 2/3,
p = 0.75,
q = 0.9,
eta = 0.25,
r_pp = 4,
kappa = 0.005,
lambda = 2+q-n,
alpha = 0.6,
ks = 4,
z0pre = 0.6,
h = 30,
beta = 100,
sigma = 1.3,
f0 = 0.5,
gamma = NA,
knife_edge_size = 1000,
gear_names = "knife_edge_gear",
...){
# If not supplied, calculate gamma using equation 2.1 in A&P 2010
if(is.na(gamma)){
alpha_e <- sqrt(2*pi) * sigma * beta^(lambda-2) * exp((lambda-2)^2 * sigma^2 / 2) # see A&P 2009
gamma <- h * f0 / (alpha_e * kappa * (1-f0)) # see A&P 2009
}
w_inf <- 10^seq(from=log10(min_w_inf), to = log10(max_w_inf), length=no_sp)
w_mat <- w_inf * eta
# Check gears
if (length(knife_edge_size) > no_sp){
stop("There cannot be more gears than species in the model")
}
if ((length(knife_edge_size) > 1) & (length(knife_edge_size) != no_sp)){
warning("Number of gears is less than number of species so gear information is being recycled. Is this what you want?")
}
if ((length(gear_names) != 1) & (length(gear_names) != no_sp)){
stop("Length of gear_names argument must equal the number of species.")
}
# Make the species parameters data.frame
trait_params_df <- data.frame(
species = 1:no_sp,
w_inf = w_inf,
w_mat = w_mat,
h = h, # max food intake
gamma = gamma, # vol. search rate,
ks = ks,# standard metabolism coefficient,
beta = beta,
sigma = sigma,
z0 = z0pre * w_inf^(n-1), # background mortality
alpha = alpha,
#r_max = r_max,
sel_func = "knife_edge",
knife_edge_size = knife_edge_size,
gear = gear_names,
erepro = 1 # not used but included out of necessity
)
# Make the therMizerParams
trait_params <- therMizerParams(trait_params_df, min_w = min_w, max_w=max_w, no_w = no_w, min_w_pp = min_w_pp, w_pp_cutoff = w_pp_cutoff, n = n, p=p, q=q, r_pp=r_pp, kappa=kappa, lambda = lambda)
# Sort out maximum recruitment - see A&P 2009
# Get max flux at recruitment boundary, R_max
# R -> | -> g0 N0
# R is egg flux, in numbers per time
# Actual flux at recruitment boundary = RDD = NDD * g0 (where g0 is growth rate)
# So in our BH SRR we need R_max comparable to RDI (to get RDD)
# R_max = N0_max * g0 (g0 is the average growth rate of smallest size, i.e. at f0 = 0.5)
# N0 given by Appendix A of A&P 2010 - see Ken's email 12/08/13
# Taken from Ken's code 12/08/13 - equation in paper is wrong!
alpha_p <- f0 * h * beta^(2 * n - q - 1) * exp((2 * n * (q - 1) - q^2 + 1) * sigma^2 / 2)
alpha_rec <- alpha_p / (alpha * h * f0 - ks)
# Calculating dw using Ken's code - see Ken's email 12/08/13
tmpA <- w_inf[1]
tmpB <- (log10(w_inf[length(w_inf)]) - log10(w_inf[1])) / (no_sp - 1) # Difference between logged w_infs, fine
dw_winf <- tmpB * tmpA *10^(tmpB*((1:no_sp)-1)) # ?
N0_max <- k0 * w_inf^(n*2-q-3+alpha_rec) * dw_winf # Why * dw_winf, not / ? Ken confirms * in email
# No need to include (1 - psi) in growth equation because allocation to reproduction at this size = 0, so 1 - psi = 1
g0 <- (alpha * f0 * h * trait_params@w[1]^n - ks * trait_params@w[1]^p)
r_max <- N0_max * g0
trait_params@species_params$r_max <- r_max
return(trait_params)
}
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