untb: Ecological Drift under the UNTB

caruso <-
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0L, 27L), .Dim = c(194L, 5L), .Dimnames = list(c("Hypochthonius_luteus", 
"Eniochthonius_minutissimus", "Sphaerochthonius_splendidus", 
"Brachychthonius_hirtus", "Brachychthonius_impressus", "Brachychthonius_jacoti", 
"Brachychthonius_pius", "Liochthonius_brevis", "Liochthonius_hystricinus", 
"Liochthonius_horridus", "Liochthonius_leptaleus", "Liochthonius_muscorum", 
"Liochthonius_neglectus", "Liochthonius_sellnicki", "Liochthonius_simplex", 
"Liochthonius_strenzkei", "Poecilochthonius_italicus", "Sellnickochthonius_cricoides", 
"Sellnickochthonius_immaculatus", "Sellnickochthonius_meridionalis", 
"Sellnickochthonius_muscorum", "Sellnickochthonius_rostratus", 
"Sellnickochthonius_suecicus", "Sellnickochthonius_zelawaiensis", 
"Synchthonius_crenulatus", "Synchthonius_elegans", "Paraliochthonius_globulifer", 
"Paraliochthonius_occultus", "Phthiracarus_anonymus", "Phthiracarus_globosus", 
"Phthiracarus_italicus", "Phthiracarus_laevigatus", "Phthiracarus_nitens", 
"AtropacarusA_clavigerus", "AtropacarusA_plakatisi", "Atropacarus_striculus", 
"SteganacarusS_magnus", "Euphthiracarus_monodactylus", "Paratritia_baloghi", 
"Rysotritia_ardua", "Eulohmannia_ribagai", "EpilohmanniaE_cylindrica", 
"Epilohmannia_schusteri", "Nothrus_anauniensis", "Camisia_horrida", 
"Camisia_segnis", "Camisia_spinifer", "Nanhermannia_nana", "Hermanniella_dolosa", 
"Hermanniella_picea", "Arthrodamaeus_mediterraneus", "Lopheremaeus_laminipes", 
"Licnodamaeus_pulcherrimus", "Licnodamaeus_undulatus", "Licnobelba_latiflabellata", 
"DamaeusD_riparius", "DamaeusS_verticillipes", "Metabelba_platynota", 
"Metabelbella_interlamellaris", "Neobelbacf_pseudopapillipes", 
"Porobelba_spinosa", "Cepheus_cepheiformis", "Microzetes_castrii", 
"Microzetes_mirandus", "Amerobelba_decedens", "Berndamerus_bicostatus", 
"Damaeolus_asperatus", "Damaeolus_bregetovae", "Fosseremus_laciniatus", 
"Amerus_cuspidatus", "Amerus_troisi", "Eueremaeus_silvestris", 
"Microzetorchestes_emeryi", "Zetorcheses_falzonii", "Dorycranosus_acutus", 
"Liacarus_brevilamellatus", "Liacarus_coracinus", "Liacarus_nitens", 
"Liacarus_subterraneus", "Liacarus_xylariae", "Xenillus_clypeator", 
"Xenillus_tegeocranus", "Cultroribula_bicultrata", "Ceratoppia_bipilis", 
"Ceratoppia_quadridentata", "Gustavia_fusifer", "Carabodes_affinis", 
"Carabodes_grandjeani", "Carabodes_marginatus", "Carabodes_minusculus", 
"Carabodes_quadrangulus", "Tectocepheus_sarekensis", "Tectocepheus_velatus", 
"Raphigneta_numidiana", "Machuella_draconis", "Anomaloppia_sp1", 
"Anomaloppia_sp2", "Berniniella_bicarinata", "Berniniellacf_hauseri", 
"DiscoppiaC_cylindrica", "Dissorhina_ornata", "Hypogeoppia_hypogea", 
"Hypogeoppia_terricola", "Hypogeoppia_sp", "Lauroppia_falcata", 
"Lauroppia_fallax", "Lauroppia_similifallax", "Lauroppia_tenuipectinata", 
"Medioppiacf_obsoleta", "Medioppiacf_subpectinata", "Medioppia_sp1", 
"Microppia_minus", "Multioppiacf_glabra", "Multioppiacf_moritzi", 
"Multioppiacf_wilsoni", "NeotrichoppiaC_confinis", "Oppia_arcidiaconae", 
"Oppia_denticulata", "OppiellaO_nova", "Oxyoppioides_decipiens", 
"RamusellaI_insculpta", "RamusellaR_assimilis", "RamusellaR_strinatii", 
"Serratoppia_serrata", "Serratoppia_sp", "Quadroppiacf_micaeli", 
"Quadroppiacf_quadricarinata", "Quadroppiacf_pseudocircumita", 
"Quadroppiacf_ritza", "Suctobelbacf_lapidaria", "Suctobelbacf_regia", 
"Suctobelbacf_trigona", "Suctobelbella_sp", "Oribella_pectinata", 
"Banksinomidae_newsp", "Trizetes_pyramidalis", "Scapheremaeus_patella", 
"Micreremus_brevipes", "Liebstadia_badensis", "Protoribates_capucinus", 
"Protoribates_lophotrichus", "Dometorina_plantivaga", "Oribatula_caliptera", 
"Oribatula_tibialis", "Zygoribatula_exarata", "Zygoribatula_exilis", 
"Haplozetes_tenuifusus", "Haplozetes_vindobonensis", "Hemileius_initialis", 
"Scheloribates_laevigatus", "Scheloribates_latipes", "Scheloribatescf_latipes", 
"Scheloribates_pallidulus", "Scheloribates_quintus", "Scheloribates_xylobatoides", 
"Scheloribates_sp1", "Scheloribates_sp2", "Chamobates_borealis", 
"Xyphobates_sp", "Euzetes_globulus", "Ceratozetes_conjuctus", 
"Ceratozetes_gracilis", "Ceratozetescf_laticuspidatus", "Ceratozetes_mediocris", 
"Ceratozetes_peritus", "Latilamellobates_incisellus", "Trichoribates_angustatus", 
"Minunthozetes_pseudofusiger", "Punctoribates_punctum", "Humerobates_rostrolamellatus", 
"Eupelops_acromios", "Eupelops_curtipilus", "Eupelops_nepotulus", 
"Eupelops_subexutus", "Eupelops_torulosus", "Peloptulus_phaeonotus", 
"Joelia_fiorii", "Ophidiotrichus_connexus", "Oribatella_brevicuspidis", 
"Oribatella_hungarica", "Oribatella_ornata", "Oribatella_pulchra", 
"Oribatella_superbula", "Achipteria_coleoptrata", "Achipteria_nitens", 
"Parachipteria_punctata", "Acrogalumna_longipluma", "Alloalumna_alamellae", 
"Galumna_europaea", "Galumna_lanceata", "Galumna_tarsipennata", 
"Pergaluma_myrmophila", "Pilogalumna_crassiclava", "Pilogalumna_tenuiclava"
), c("Beech", "Coppice", "Grassland", "Heatland", "Biancana")))