Description Usage Arguments Details Value Warnings Note References See Also Examples
This function estimates the probability of occurrence using presence-only data and spatially-referenced covariates. Species distribution maps can be created by plotting the expected values of occurrence probability. The model is described by Royle et al. (2012).
1 2 3 4 |
formula |
A right-hand side |
rasters |
The spatially-referenced covariate data formatted as a 'raster
stack' created by the |
points |
A |
x |
A |
z |
A |
link |
The link function. Either "logit" (the default) or "cloglog". |
starts |
Starting values for the parameters. This should be a vector with as many elements as there are parameters. By default, all starting values are 0, which should be adequate if covariates are standardized. |
hessian |
Logical. Should the hessian be computed and the variance-covariance matrix returned? |
fixed |
Optional vector for fixing parameters. It must be
of length equal to the number of parameters in the
model. If an element of |
removeDuplicates |
Logical. Should duplicate points be removed? Defaults to FALSE, but note that the MAXENT default is TRUE. |
savedata |
Should the raster data be saved with the fitted model? Defaults to FALSE in order to reduce the size of the returned object. If you wish to make predictions, it is safer to set this to TRUE, otherwise the raster data are searched for in the working directory, and thus may not be the data used to fit the model. |
na.action |
See |
... |
Additional arguments passed to |
points
and rasters
should the same coordinate system. The program does not check
this so it is up to the user.
A list with 8 components
Est |
data.frame containing the parameter estimates (Ests) and standard errors (SE). |
vcov |
variance-covariance matrix |
AIC |
AIC |
call |
the original call |
pts.removed |
The points removed due to missing values |
pix.removed |
The pixels removed due to missing values |
optim |
The object returned by |
not.fixed |
A logical vector indicating if a parameter was estimated or fixed. |
link |
The link function |
Maximizing the log-likelihood function is achieved using the
optim
function, which can fail to find the global optima
if sensible starting values are not
supplied. The default starting values are rep(0, npars)
, which
will often be adequate if the covariates have been
standardized. Standardizing covariates is thus recommended.
Even when covariates are standardized, it is always a good idea to try
various starting values to see if the
log-likelihood can be increased. When fitting models with many
parameters, good starting values can be found by fitting simpler
models first.
In general it is very hard to obtain a random sample of presence
points, which is a requirement of both the Royle et al. (2012)
method and of MAXENT. This is one of many reasons why presence-absence
data are preferable to presence-only data. When presence-absence data
are available, they can be modeled using functions such as
glm
. Creating species distribution maps
from glm
is easily accomplished using the
predict
method.
The MAXENT software assumes that species prevalence is known a
priori. If the user does not specify a value for prevalence,
prevalence is set to 0.5. MAXENT predictions of occurrence probability
are highly sensitive to this setting. In contrast, maxlike
directly estimates prevalence.
Another weakness of models for presence-only data is that they do not allow one to model detection probability, which is typically less than one in field conditions. If detection probability is affected by the same covariates that affect occurrence probability, then bias is inevitable. The R package unmarked (Fiske and Chandler 2011) offers numerous methods for jointly modeling both occurrence and detection probability when detection/non-detection data are available.
Royle, J.A., R.B. Chandler, C. Yackulic and J. D. Nichols. 2012. Likelihood analysis of species occurrence probability from presence-only data for modelling species distributions. Methods in Ecology and Evolution. doi: 10.1111/j.2041-210X.2011.00182.x
Fiske, I. and R.B. Chandler. 2011. unmarked: An R Package for Fitting Hierarchical Models of Wildlife Occurrence and Abundance. Journal of Statistical Software 43(10).
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 74 75 76 77 78 79 80 81 82 83 84 85 86 87 88 89 90 91 92 93 94 95 96 97 98 99 100 101 102 103 104 105 106 107 108 109 110 111 112 113 114 115 116 | # Carolina Wren data used in Royle et. al (2012)
data(carw)
# Covert data.frame to a list of rasters
rl <- lapply(carw.data$raster.data, function(x) {
m <- matrix(x, nrow=carw.data$dim[1], ncol=carw.data$dim[2], byrow=TRUE)
r <- raster(m)
extent(r) <- carw.data$ext
r
})
# Create a raster stack and add layer names
rs <- stack(rl[[1]], rl[[2]], rl[[3]], rl[[4]], rl[[5]], rl[[6]])
names(rs) <- names(carw.data$raster.data)
plot(rs)
# Fit a model
fm <- maxlike(~pcMix + I(pcMix^2) + pcDec + I(pcDec^2)+ pcCon +
I(pcCon^2) + pcGr + I(pcGr^2) +
Lat + I(Lat^2) + Lon + I(Lon^2), rs, carw.data$xy1,
method="BFGS", removeDuplicates=TRUE, savedata=TRUE)
summary(fm)
confint(fm)
AIC(fm)
logLik(fm)
# Produce species distribution map (ie, expected probability of occurrence)
psi.hat <- predict(fm) # Will warn if savedata=FALSE
plot(psi.hat)
points(carw.data$xy1, pch=16, cex=0.1)
# MAXENT sets "default prevalence" to an arbitrary value, 0.5.
# We could do something similar by fixing the intercept at logit(0.5)=0.
# However, it seems more appropriate to estimate this parameter.
fm.fix <- update(fm, fixed=c(0, rep(NA,length(coef(fm))-1)))
# Predict data.frame
presenceData <- as.data.frame(extract(rs, carw.data$xy1))
presenceData <- presenceData[complete.cases(presenceData), ]
presence.predictions <- predict(fm, newdata=presenceData)
summary(presence.predictions)
# Calibrate with data.frames
PresenceUniqueCells <- unique(cellFromXY(rs, xy=carw.data$xy1))
PresenceUnique <- xyFromCell(rs, PresenceUniqueCells)
presenceData <- as.data.frame(extract(rs, PresenceUnique))
library(dismo)
background <- randomPoints(rs, n=ncell(rs), extf=1.00)
backgroundData <- as.data.frame(extract(rs, y=background))
backgroundData <- backgroundData[complete.cases(backgroundData), ]
fm2 <- maxlike(~pcMix + I(pcMix^2) + pcDec + I(pcDec^2)+ pcCon +
I(pcCon^2) + pcGr + I(pcGr^2) +
Lat + I(Lat^2) + Lon + I(Lon^2),
rasters=NULL, points=NULL,
x=presenceData, z=backgroundData,
method="BFGS", removeDuplicates=TRUE, savedata=TRUE)
summary(fm2)
fm2$rasters <- rs
psi.hat2 <- predict(fm2)
## Not run:
# Simulation example
set.seed(131)
x1 <- sort(rnorm(100))
x1 <- raster(outer(x1, x1), xmn=0, xmx=100, ymn=0, ymx=100)
x2 <- raster(matrix(runif(1e4), 100, 100), 0, 100, 0, 100)
# Code factors as dummy variables.
# Note, using asFactor(x3) will not help
x3 <- raster(matrix(c(0,1), 100, 100), 0, 100, 0, 100)
logit.psi <- -1 + 1*x1 + 0*x2
psi <- exp(logit.psi)/(1+exp(logit.psi))
plot(psi)
r <- stack(x1, x2, x3)
names(r) <- c("x1", "x2", "x3")
plot(r)
pa <- matrix(NA, 100, 100)
pa[] <- rbinom(1e4, 1, as.matrix(psi))
str(pa)
table(pa)
pa <- raster(pa, 0, 100, 0, 100)
plot(pa)
xy <- xyFromCell(pa, sample(Which(pa==1, cells=TRUE), 1000))
plot(x1)
points(xy)
fm2 <- maxlike(~x1 + x2 + x3, r, xy)
summary(fm2)
confint(fm2)
AIC(fm2)
logLik(fm2)
## End(Not run)
|
Add the following code to your website.
For more information on customizing the embed code, read Embedding Snippets.