nichevar <-
function (P, D = NULL, q = NULL, mode="multiple", Np = NULL, Nq = NULL, nboot = 1000, alpha=0.05) {
if (!inherits(P, "data.frame")) stop("Non convenient dataframe for species resource use")
if (!is.null(D)) {
if (!inherits(D, "dist")) stop("Object of class 'dist' expected for distance")
D <- as.matrix(D)
if (ncol(P) != nrow(D)) stop("The number of columns in P must be equal to the number of items in D")
D <- as.dist(D)
}
if(!is.null(Np) && mode=="multiple") {
if(length(Np)!=nrow(P)) stop("The number of items in Np must be equal to the number of rows in P")
}
if(!is.null(q)) {
if(length(q)!=ncol(P)) stop("The number of items in q must be equal to the number of columns in P")
q = q/sum(q) #Just to check that we have proportions
} else {
#If no availability is provided, then all resources are equally available
q = rep(1/ncol(P),ncol(P))
}
#If no distance matrix is provided, the distance between resources is assumed to be maximum
if (is.null(D)) D <- as.dist((matrix(1, ncol(P), ncol(P)) - diag(rep(1, ncol(P)))))
# Computes the niche breadth from the resource preferences of the target and the resource relationships
nichevar1<-function(f, D) {
if (is.na(sum(f))) v <- NA
else if (sum(f) < 1e-16) v <- 0
else v <- (f %*% (as.matrix(D)^2) %*% f)/(2*(sum(f)^2))
return(v)
}
# Returns preference from a resource use vector (considering resource availability in desired)
getF<-function(p,q=NULL) {
if(!is.null(q)) {
a = p/q
return(a/sum(a))
} else { #Just to check that we have proportions
return(p/sum(p))
}
}
if(!is.null(Np) || mode=="single") nc = 3
else nc = 1
#Rows in P are different niches
if(mode=="multiple") {
B <- as.data.frame(matrix(0,nrow=nrow(P), ncol=nc))
rownames(B) <- row.names(P)
for (i in 1:nrow(P)) {
pi = as.numeric(P[i,])
B[i,1] = nichevar1(getF(pi,q), D)
if(!is.null(Np)) {
BB = vector("numeric",length=nboot)
if(sum(is.na(getF(pi)))==0) {
#Generate bootstrap samples from multinomial distribution
psamp = rmultinom(nboot,Np[i],getF(pi))
if(!is.null(Nq)) qsamp = rmultinom(nboot,Nq,q)
for(b in 1:nboot) {
if(!is.null(Nq)) BB[b] = nichevar1(getF(psamp[,b],qsamp[,b]),D)
else BB[b] = nichevar1(getF(psamp[,b],q),D)
}
#Some NA may appear because of zeroes in qsamp
BB = BB[!is.na(BB)]
#Compute Bias-corrected percentile method (Manly 2007: pp52-56)
z0 = qnorm(sum(BB<B[i,1])/length(BB))
lj = floor(length(BB)*pnorm(2*z0+qnorm(alpha/2)))
uj = floor(length(BB)*pnorm(2*z0+qnorm(1-(alpha/2))))
if(lj > 0 && uj > 0) {
sbb = sort(BB)
B[i,2] = sbb[lj]
B[i,3] = sbb[uj]
}
}
}
}
}
#Rows in P are observations
else if(mode=="single") {
B <- as.data.frame(matrix(0,nrow=1, ncol=nc))
rownames(B) <- "Niche"
B[1,1] = nichevar1(getF(colSums(P),q), D)
BB = vector("numeric",length=nboot)
if(!is.null(Nq)) qsamp = rmultinom(nboot,Nq,q)
for(b in 1:nboot) {
psamp = colSums(P[sample(1:nrow(P),replace=TRUE),])
if(!is.null(Nq)) {
BB[b] = nichevar1(getF(psamp,qsamp[b]),D)
} else {
BB[b] = nichevar1(getF(psamp,q),D)
}
}
#Some NA may appear because of zeroes in qsamp
BB = BB[!is.na(BB)]
#Compute Bias-corrected percentile method (Manly 2007: pp52-56)
z0 = qnorm(sum(BB<B[1,1])/length(BB))
lj = floor(length(BB)*pnorm(2*z0+qnorm(alpha/2)))
uj = floor(length(BB)*pnorm(2*z0+qnorm(1-(alpha/2))))
if(lj > 0 && uj > 0) {
sbb = sort(BB)
B[1,2] = sbb[lj]
B[1,3] = sbb[uj]
}
}
if(nc==1) names(B) <- "B"
else names(B) <- c("B","LC", "UC")
return(B)
}
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