R/runpoll_3seasons.R
In yclough/poll4pop: Functions for modelling central-place foraging pollinators
Defines functions
runpoll_3seasons
Documented in
runpoll_3seasons
#' Pollinator model function
#'
#' Uses spatial data on floral and nesting quality to compute flower visitation and population dynamics for wild pollinators for a given year.
#' The function can be used for mutliple years by running a loop and using as input the output of the previous model (see example).
#' @param M_poll0 the initial population sizes to be used only if we are not in the first year
#' @param firstyear a boolean identifying whether we run the model for the first time (in which case we need to initialize the model)
#' @param firstyearfactor a factor to control the initial population sizes, defaults to 1.
#' @param bees a vector of bees identifiers corresponding to those in the parameter files
#' @param cell.size the size of a cell in the landscape matrices, in meters (default to 25)
#' @param paramList the list of parameters needed to run the model
#' @param nest the rasterstack containing the nesting qualities (dimensions should be consistent with the number of bees in the model)
#' @param floral the rasterstack containing the floral values (dimensions should be consistent with the number of bees in the model)
#' @param cutoff, the percentile at which the dispersal kernel is cut off
#' @param loc_managed, the location at which managed pollinators are placed (e.g. apiaries)
#' @return a list containing :
#' \describe{
#' \item{M_poll}{the number of queens at the end of each period and at the beginning of the next season}
#' \item{N_poll}{the number of foraging bees in each period}
#' \item{flowvis}{the visitation rates per cell in each period}
#' \item{floral and nest}{the floral and nesting values (the same as in the inputs, but stored in the results for convenience)}
#' \item{pollres}{the resources gathered by the bees}
#' }
#'
#' @references Haeussler J, Sahlin U, Baey C, Smith HG, Clough Y (2017) Predicting pollinator population size and pollination ecosystem service responses
#' to enhancing floral and nesting resources. Ecology and Evolution, 7: 1898-1908.\url{http://dx.doi.org/10.1002/ece3.2765}
#' Expansion to 3 seasons first used in:
#' Gardner E, et al. (2020) Reliably predicting pollinator abundance: Challenges of calibrating
#' process-based ecological models. Methods in Ecology and Evolution. \url{}
#'
#' @export
#'
#' @examples
#'
#' nf<-computeFloralNesting(landuseMap=landuse, edgesMap=stack(grassmargins,flowermargins), unitEdges = "sqm", widthEdges=1,
#' landuseCodes, bees=c("GroundNestingBumblebees", "GroundNestingSolitaryBees"), num_floral=3,
#' florNestInfo=parameters$florNestInfo, codeEdges=c(11,21), cell.size = 10,paramList=parameters)
#' poll<-runpoll_3seasons(M_poll0 = numeric(0), firstyear=TRUE, firstyearfactor = c(1, 1),
#' bees = c("GroundNestingBumblebees", "GroundNestingSolitaryBees"), cell.size = 10, paramList=parameters, nest=nf$nest,
#' floral=nf$floral, cutoff = 0.99, loc_managed)
runpoll_3seasons<-function(M_poll0=numeric(0),
firstyear,
firstyearfactor=c(1,1),
bees = c("Bombus spp.","Solitary spp."),
cell.size=25,
paramList,
nest,
floral,
cutoff=0.99,
loc_managed){
require(raster)
nr <- nrow(nest[[1]])
nc <- ncol(nest[[1]])
ncells <- ncell(nest[[1]])
hab_names<-as.character(unique(paramList$lfn[,'lu']))
bee_names<-paramList$poll_names$species_name
bsel <- match(bees,as.character(paramList$poll_names$species_name))
num_hab<-length(unique(paramList$lfn[,'code']))
num_bees=length(bsel)
num_floral<- nlayers(floral[[1]]) # takes the info from the first species
# proportion of foraging workers
pw <- paramList$growth[paramList$growth$Short.notation=='pw',][bsel,3]
av<-array(paramList$av[ bsel,'best.guess'],c(num_bees))
solitary_social<-paramList$poll_names$solitary_social[bsel]
wild_managed<-paramList$poll_names$wild_managed[bsel]
foraging_period <- paramList$poll_names[bsel,c('foraging_P1','foraging_P2','foraging_P3')] #Edited!!!
growth <- array(0,c(3,2,num_bees)) #One column for q and one for w
# Derive growth parameters for the different species #
for(s in 1:num_bees){
# identify subset of growth parameters for s
selec=which(paramList$growth$species==bsel[s])
w=1 # first period, i.e. queens
ind<-c(grep("aq",paramList$gr[selec,'Short.notation']),grep("bq",paramList$gr[selec,'Short.notation']),grep("QEmax",paramList$gr[selec,'Short.notation']))
growth[,w,s]<-paramList$gr[selec,][ind,3] #queens
w=2
ind<-c(grep("aw",paramList$gr[selec,'Short.notation']),grep("bw",paramList$gr[selec,'Short.notation']),grep("Wmax",paramList$gr[selec,'Short.notation']))
growth[,w,s]<-paramList$gr[selec,][ind,3] #workers
}
# Define pollinator population size, flower visitation and pollinator resource storage arrays #
emptyraster<-nest[[1]]
values(emptyraster)<-NA
# M_poll is the number of queens and N_poll the number of foragers (e.g. workers)
M_poll <- stack(mget(rep("emptyraster",num_floral+1)))
names(M_poll)=paste("period",1:(num_floral+1),sep=" ")
M_poll <- mget(rep("M_poll",num_bees))
names(M_poll)<-bees
N_poll <- stack(mget(rep("emptyraster",num_floral)))
names(N_poll)=paste("period",1:(num_floral),sep=" ")
N_poll <- mget(rep("N_poll",num_bees))
names(N_poll)<-bees
floral_periods <- stack(mget(rep("emptyraster",num_floral)))
names(floral_periods)=paste("floral period",1:num_floral,sep=" ")
flowvis <- mget(rep("floral_periods",num_bees))
names(flowvis)=bees
pollres<-mget(rep("floral_periods",num_bees)) # resource availability in the surrounding landscape for pollinators nesting in the focal cell
names(pollres)=bees
### ------------------------------------------###
### Kernel computation ###
### ------------------------------------------###
##print("Kernel computation ...")
# The kernels are computed at the beginning and used throughout the simulation
kernelFor <- list()
kernelNest <- list()
for (s in 1:num_bees)
{
kernelFor[[s]] <- kerncalc(
paramList$distance[paramList$distance$species==bsel[s]¶mList$distance$activity=="foraging","best_guess"], cell.size, maxSize = floor(min((nr-1)/2,(nc-1)/2)), decaycut=cutoff)$decay
kernelNest[[s]] <- kerncalc(
paramList$distance[paramList$distance$species==bsel[s]¶mList$distance$activity=="nesting","best_guess"], cell.size, maxSize = floor(min((nr-1)/2,(nc-1)/2)), decaycut=cutoff)$decay
}
# Initialize population size (1st year) #
if(firstyear)
{
##print("Initiation ...")
for(s in 1:num_bees)
{
if(wild_managed[s]=="wild"){
values(M_poll[[s]][[1]]) <- firstyearfactor[s]*values(nest[[s]])*av[s]*cell.size^2/(100^2)
#print(mean(values(M_poll[[s]][[1]])))
values(M_poll[[s]][[1]]) <- rpois(length(values(M_poll[[s]][[1]])), values(M_poll[[s]][[1]]))
}
if(wild_managed[s]=="managed"){
values(M_poll[[s]][[1]]) <- values(loc_managed)
}
}
}else{
for(s in 1:num_bees)
{
values(M_poll[[s]][[1]])<-values(M_poll0[[s]][[(num_floral+1)]]) #Edited!!!
}
}
for(s in 1:num_bees)
{
##print(bees[s]);flush.console()
##print(paste("nb females start : ", round(sum(values(M_poll[[s]][[1]]))) ));flush.console()
## ----------------- floral period 1 ----------------- ##
v=1
##print("Floral period 1 ...");flush.console()
## M_poll (number of queens) aggregate resources (pollres v=1) --> VR1
# for all species foraging in period 1
if(foraging_period[s,1]!="none")
{
foraging <- latfordisp(N=raster::as.matrix(M_poll[[s]][[1]]),
alpha = raster::as.matrix(floral[[s]][[v]]),
decay = kernelFor[[s]])
values(flowvis[[s]][[v]]) <- as.vector(t(foraging$visitation_rate)) # flower visitation in the cells surrounding the focal cell
values(pollres[[s]][[v]]) <- as.vector(t(foraging$Racc_per_forager)) # aggregated resources experienced by the pollinator nesting in the focal cell
}
# for social species with queens foraging in period 1
if(solitary_social[s]=="social" & foraging_period[s,v]=="q")
{
## ------ # Population growth WORKERS # ------- ##
## N_poll, number of workers produced, depending on the number of queens and resources in period 1
values(N_poll[[s]][[1]]) <- values(M_poll[[s]][[1]]) * growth.func(R = values(pollres[[s]][[v]]),
a = growth[1,2,s],
b = growth[2,2,s],
max = growth[3,2,s])
##print("growth func ran ok");flush.console()
}
# for solitary species foraging in period 1
if(solitary_social[s]=="solitary" & foraging_period[s,v]=="q")
{
values(M_poll[[s]][[2]]) <- values(M_poll[[s]][[1]]) * growth.func(R = values(pollres[[s]][[v]]),
a = growth[1,1,s],
b = growth[2,1,s],
max = growth[3,1,s])
M_pollend = M_poll[[s]][[2]] #Added!!!
}
## ----------------- floral period 2 ----------------- ##
v=2
##print("Floral period 2 ...");flush.console()
if(solitary_social[s]=="social" & foraging_period[s,v]=="w")
{
## N_poll (workers) aggregate resources (pollres v=2) --> VR2
## Only a proportion pw of the workers is actually foraging
foraging <- latfordisp(N = pw[s]*raster::as.matrix(N_poll[[s]][[1]]), alpha = raster::as.matrix(floral[[s]][[v]]), decay = kernelFor[[s]])
values(flowvis[[s]][[v]]) <- as.vector(t(foraging$visitation_rate)) # flower visitation in the cells surrounding the focal cell
values(pollres[[s]][[v]]) <- as.vector(t(foraging$Racc_per_forager)) #
## ------ # Population growth QUEENS # ------- ##
## M_poll produced at the end of the season depending on M_poll and resources in period 2
# growth[3,2,s] is the maximal number of workers
# N_poll is the number of foraging bees
#values(M_poll[[s]][[2]]) <- values(M_poll[[s]][[1]]) * growth.func(R = values(pollres[[s]][[v]])*pw[s]*values(N_poll[[s]][[1]])/values(M_poll[[s]][[1]]), a = growth[1,1,s], b = growth[2,1,s], max = growth[3,1,s]) #Replaced!!!
M_poll[[s]][[2]] = M_poll[[s]][[1]] #Added!!!
values(N_poll[[s]][[2]]) <- values(N_poll[[s]][[1]])+values(M_poll[[s]][[1]]) * growth.func(R = values(pollres[[s]][[v]])*pw[s]*values(N_poll[[s]][[1]])/values(M_poll[[s]][[1]]),
a = growth[1,2,s],
b = growth[2,2,s],
max = growth[3,2,s]) #Added!!!
}
if(solitary_social[s]=="solitary" & foraging_period[s,v]=="q")
{
foraging <- latfordisp(N=raster::as.matrix(M_poll[[s]][[1]]),
alpha = raster::as.matrix(floral[[s]][[v]]),
decay = kernelFor[[s]])
values(flowvis[[s]][[v]]) <- as.vector(t(foraging$visitation_rate)) # flower visitation in the cells surrounding the focal cell
values(pollres[[s]][[v]]) <- as.vector(t(foraging$Racc_per_forager)) # aggregated resources experienced by the pollinator nesting in the focal cell
values(M_poll[[s]][[2]]) <- values(M_poll[[s]][[1]]) #Added!!!
values(M_poll[[s]][[3]]) <- values(M_poll[[s]][[2]]) * growth.func(R = values(pollres[[s]][[v]]),
a = growth[1,1,s],
b = growth[2,1,s],
max = growth[3,1,s]) #Edited!!!
values(M_poll[[s]][[1]]) <- NA #Added!!!
M_pollend = M_poll[[s]][[3]] #Added!!!
}
## ----------------- floral period 3 ----------------- ##Added!!!
v=3
##print("Floral period 3 ...");flush.console()
if(solitary_social[s]=="social" & foraging_period[s,v]=="w")
{
## N_poll (workers) aggregate resources (pollres v=2) --> VR2
## Only a proportion pw of the workers is actually foraging
foraging <- latfordisp(N = pw[s]*raster::as.matrix(N_poll[[s]][[2]]), alpha = raster::as.matrix(floral[[s]][[v]]), decay = kernelFor[[s]])
values(flowvis[[s]][[v]]) <- as.vector(t(foraging$visitation_rate)) # flower visitation in the cells surrounding the focal cell
values(pollres[[s]][[v]]) <- as.vector(t(foraging$Racc_per_forager)) #
## ------ # Population growth QUEENS # ------- ##
## M_poll produced at the end of the season depending on M_poll and resources in period 2
# growth[3,2,s] is the maximal number of workers
# N_poll is the number of foraging bees
values(M_poll[[s]][[3]]) <- values(M_poll[[s]][[1]]) * growth.func(R = values(pollres[[s]][[v]])*pw[s]*values(N_poll[[s]][[2]])/values(M_poll[[s]][[1]]), a = growth[1,1,s], b = growth[2,1,s], max = growth[3,1,s]) #Edited!!!
M_pollend = M_poll[[s]][[3]] #Added!!!
##print("growth func ran ok");flush.console()
}
if(solitary_social[s]=="solitary" & foraging_period[s,v]=="q")
{
print("Don't allow Season 3 only active solitary bees") #Added!!!
}
##print(paste("nb new females : ", round(sum(values(M_pollend))) ));flush.console() #Edited!!!
## M_poll at the beginning of the next season depending on winter survival of M_poll at the end of the season
# non-honeybees disperse to their new nesting sites #
if(wild_managed[s]=="wild")
{
poll.out.start <- latfordisp(N = raster::as.matrix(M_pollend),
alpha = raster::as.matrix(nest[[s]]),
decay = kernelNest[[s]]) #Edited!!!
values(M_poll[[s]][[(num_floral+1)]]) <- pmin(as.vector(t(poll.out.start$visitation_rate)),(av[s]*cell.size^2/10000)*values(nest[[s]])) #Edited!!!
##print(paste("nb females surv : ", round(sum(values(M_poll[[s]][[(num_floral+1)]]))) ));flush.console() #Edited!!!
}
} #end species
return(list(M_poll=M_poll,
N_poll=N_poll,
flowvis=flowvis,
floral=floral,
nest=nest,
pollres=pollres))
}
yclough/poll4pop documentation built on Sept. 7, 2020, 1:13 p.m.
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Defines functions runpoll_3seasons
Documented in runpoll_3seasons
#' Pollinator model function
#'
#' Uses spatial data on floral and nesting quality to compute flower visitation and population dynamics for wild pollinators for a given year.
#' The function can be used for mutliple years by running a loop and using as input the output of the previous model (see example).
#' @param M_poll0 the initial population sizes to be used only if we are not in the first year
#' @param firstyear a boolean identifying whether we run the model for the first time (in which case we need to initialize the model)
#' @param firstyearfactor a factor to control the initial population sizes, defaults to 1.
#' @param bees a vector of bees identifiers corresponding to those in the parameter files
#' @param cell.size the size of a cell in the landscape matrices, in meters (default to 25)
#' @param paramList the list of parameters needed to run the model
#' @param nest the rasterstack containing the nesting qualities (dimensions should be consistent with the number of bees in the model)
#' @param floral the rasterstack containing the floral values (dimensions should be consistent with the number of bees in the model)
#' @param cutoff, the percentile at which the dispersal kernel is cut off
#' @param loc_managed, the location at which managed pollinators are placed (e.g. apiaries)
#' @return a list containing :
#' \describe{
#' \item{M_poll}{the number of queens at the end of each period and at the beginning of the next season}
#' \item{N_poll}{the number of foraging bees in each period}
#' \item{flowvis}{the visitation rates per cell in each period}
#' \item{floral and nest}{the floral and nesting values (the same as in the inputs, but stored in the results for convenience)}
#' \item{pollres}{the resources gathered by the bees}
#' }
#'
#' @references Haeussler J, Sahlin U, Baey C, Smith HG, Clough Y (2017) Predicting pollinator population size and pollination ecosystem service responses
#' to enhancing floral and nesting resources. Ecology and Evolution, 7: 1898-1908.\url{http://dx.doi.org/10.1002/ece3.2765}
#' Expansion to 3 seasons first used in:
#' Gardner E, et al. (2020) Reliably predicting pollinator abundance: Challenges of calibrating
#' process-based ecological models. Methods in Ecology and Evolution. \url{}
#'
#' @export
#'
#' @examples
#'
#' nf<-computeFloralNesting(landuseMap=landuse, edgesMap=stack(grassmargins,flowermargins), unitEdges = "sqm", widthEdges=1,
#' landuseCodes, bees=c("GroundNestingBumblebees", "GroundNestingSolitaryBees"), num_floral=3,
#' florNestInfo=parameters$florNestInfo, codeEdges=c(11,21), cell.size = 10,paramList=parameters)
#' poll<-runpoll_3seasons(M_poll0 = numeric(0), firstyear=TRUE, firstyearfactor = c(1, 1),
#' bees = c("GroundNestingBumblebees", "GroundNestingSolitaryBees"), cell.size = 10, paramList=parameters, nest=nf$nest,
#' floral=nf$floral, cutoff = 0.99, loc_managed)
runpoll_3seasons<-function(M_poll0=numeric(0),
firstyear,
firstyearfactor=c(1,1),
bees = c("Bombus spp.","Solitary spp."),
cell.size=25,
paramList,
nest,
floral,
cutoff=0.99,
loc_managed){
require(raster)
nr <- nrow(nest[[1]])
nc <- ncol(nest[[1]])
ncells <- ncell(nest[[1]])
hab_names<-as.character(unique(paramList$lfn[,'lu']))
bee_names<-paramList$poll_names$species_name
bsel <- match(bees,as.character(paramList$poll_names$species_name))
num_hab<-length(unique(paramList$lfn[,'code']))
num_bees=length(bsel)
num_floral<- nlayers(floral[[1]]) # takes the info from the first species
# proportion of foraging workers
pw <- paramList$growth[paramList$growth$Short.notation=='pw',][bsel,3]
av<-array(paramList$av[ bsel,'best.guess'],c(num_bees))
solitary_social<-paramList$poll_names$solitary_social[bsel]
wild_managed<-paramList$poll_names$wild_managed[bsel]
foraging_period <- paramList$poll_names[bsel,c('foraging_P1','foraging_P2','foraging_P3')] #Edited!!!
growth <- array(0,c(3,2,num_bees)) #One column for q and one for w
# Derive growth parameters for the different species #
for(s in 1:num_bees){
# identify subset of growth parameters for s
selec=which(paramList$growth$species==bsel[s])
w=1 # first period, i.e. queens
ind<-c(grep("aq",paramList$gr[selec,'Short.notation']),grep("bq",paramList$gr[selec,'Short.notation']),grep("QEmax",paramList$gr[selec,'Short.notation']))
growth[,w,s]<-paramList$gr[selec,][ind,3] #queens
w=2
ind<-c(grep("aw",paramList$gr[selec,'Short.notation']),grep("bw",paramList$gr[selec,'Short.notation']),grep("Wmax",paramList$gr[selec,'Short.notation']))
growth[,w,s]<-paramList$gr[selec,][ind,3] #workers
}
# Define pollinator population size, flower visitation and pollinator resource storage arrays #
emptyraster<-nest[[1]]
values(emptyraster)<-NA
# M_poll is the number of queens and N_poll the number of foragers (e.g. workers)
M_poll <- stack(mget(rep("emptyraster",num_floral+1)))
names(M_poll)=paste("period",1:(num_floral+1),sep=" ")
M_poll <- mget(rep("M_poll",num_bees))
names(M_poll)<-bees
N_poll <- stack(mget(rep("emptyraster",num_floral)))
names(N_poll)=paste("period",1:(num_floral),sep=" ")
N_poll <- mget(rep("N_poll",num_bees))
names(N_poll)<-bees
floral_periods <- stack(mget(rep("emptyraster",num_floral)))
names(floral_periods)=paste("floral period",1:num_floral,sep=" ")
flowvis <- mget(rep("floral_periods",num_bees))
names(flowvis)=bees
pollres<-mget(rep("floral_periods",num_bees)) # resource availability in the surrounding landscape for pollinators nesting in the focal cell
names(pollres)=bees
### ------------------------------------------###
### Kernel computation ###
### ------------------------------------------###
##print("Kernel computation ...")
# The kernels are computed at the beginning and used throughout the simulation
kernelFor <- list()
kernelNest <- list()
for (s in 1:num_bees)
{
kernelFor[[s]] <- kerncalc(
paramList$distance[paramList$distance$species==bsel[s]¶mList$distance$activity=="foraging","best_guess"], cell.size, maxSize = floor(min((nr-1)/2,(nc-1)/2)), decaycut=cutoff)$decay
kernelNest[[s]] <- kerncalc(
paramList$distance[paramList$distance$species==bsel[s]¶mList$distance$activity=="nesting","best_guess"], cell.size, maxSize = floor(min((nr-1)/2,(nc-1)/2)), decaycut=cutoff)$decay
}
# Initialize population size (1st year) #
if(firstyear)
{
##print("Initiation ...")
for(s in 1:num_bees)
{
if(wild_managed[s]=="wild"){
values(M_poll[[s]][[1]]) <- firstyearfactor[s]*values(nest[[s]])*av[s]*cell.size^2/(100^2)
#print(mean(values(M_poll[[s]][[1]])))
values(M_poll[[s]][[1]]) <- rpois(length(values(M_poll[[s]][[1]])), values(M_poll[[s]][[1]]))
}
if(wild_managed[s]=="managed"){
values(M_poll[[s]][[1]]) <- values(loc_managed)
}
}
}else{
for(s in 1:num_bees)
{
values(M_poll[[s]][[1]])<-values(M_poll0[[s]][[(num_floral+1)]]) #Edited!!!
}
}
for(s in 1:num_bees)
{
##print(bees[s]);flush.console()
##print(paste("nb females start : ", round(sum(values(M_poll[[s]][[1]]))) ));flush.console()
## ----------------- floral period 1 ----------------- ##
v=1
##print("Floral period 1 ...");flush.console()
## M_poll (number of queens) aggregate resources (pollres v=1) --> VR1
# for all species foraging in period 1
if(foraging_period[s,1]!="none")
{
foraging <- latfordisp(N=raster::as.matrix(M_poll[[s]][[1]]),
alpha = raster::as.matrix(floral[[s]][[v]]),
decay = kernelFor[[s]])
values(flowvis[[s]][[v]]) <- as.vector(t(foraging$visitation_rate)) # flower visitation in the cells surrounding the focal cell
values(pollres[[s]][[v]]) <- as.vector(t(foraging$Racc_per_forager)) # aggregated resources experienced by the pollinator nesting in the focal cell
}
# for social species with queens foraging in period 1
if(solitary_social[s]=="social" & foraging_period[s,v]=="q")
{
## ------ # Population growth WORKERS # ------- ##
## N_poll, number of workers produced, depending on the number of queens and resources in period 1
values(N_poll[[s]][[1]]) <- values(M_poll[[s]][[1]]) * growth.func(R = values(pollres[[s]][[v]]),
a = growth[1,2,s],
b = growth[2,2,s],
max = growth[3,2,s])
##print("growth func ran ok");flush.console()
}
# for solitary species foraging in period 1
if(solitary_social[s]=="solitary" & foraging_period[s,v]=="q")
{
values(M_poll[[s]][[2]]) <- values(M_poll[[s]][[1]]) * growth.func(R = values(pollres[[s]][[v]]),
a = growth[1,1,s],
b = growth[2,1,s],
max = growth[3,1,s])
M_pollend = M_poll[[s]][[2]] #Added!!!
}
## ----------------- floral period 2 ----------------- ##
v=2
##print("Floral period 2 ...");flush.console()
if(solitary_social[s]=="social" & foraging_period[s,v]=="w")
{
## N_poll (workers) aggregate resources (pollres v=2) --> VR2
## Only a proportion pw of the workers is actually foraging
foraging <- latfordisp(N = pw[s]*raster::as.matrix(N_poll[[s]][[1]]), alpha = raster::as.matrix(floral[[s]][[v]]), decay = kernelFor[[s]])
values(flowvis[[s]][[v]]) <- as.vector(t(foraging$visitation_rate)) # flower visitation in the cells surrounding the focal cell
values(pollres[[s]][[v]]) <- as.vector(t(foraging$Racc_per_forager)) #
## ------ # Population growth QUEENS # ------- ##
## M_poll produced at the end of the season depending on M_poll and resources in period 2
# growth[3,2,s] is the maximal number of workers
# N_poll is the number of foraging bees
#values(M_poll[[s]][[2]]) <- values(M_poll[[s]][[1]]) * growth.func(R = values(pollres[[s]][[v]])*pw[s]*values(N_poll[[s]][[1]])/values(M_poll[[s]][[1]]), a = growth[1,1,s], b = growth[2,1,s], max = growth[3,1,s]) #Replaced!!!
M_poll[[s]][[2]] = M_poll[[s]][[1]] #Added!!!
values(N_poll[[s]][[2]]) <- values(N_poll[[s]][[1]])+values(M_poll[[s]][[1]]) * growth.func(R = values(pollres[[s]][[v]])*pw[s]*values(N_poll[[s]][[1]])/values(M_poll[[s]][[1]]),
a = growth[1,2,s],
b = growth[2,2,s],
max = growth[3,2,s]) #Added!!!
}
if(solitary_social[s]=="solitary" & foraging_period[s,v]=="q")
{
foraging <- latfordisp(N=raster::as.matrix(M_poll[[s]][[1]]),
alpha = raster::as.matrix(floral[[s]][[v]]),
decay = kernelFor[[s]])
values(flowvis[[s]][[v]]) <- as.vector(t(foraging$visitation_rate)) # flower visitation in the cells surrounding the focal cell
values(pollres[[s]][[v]]) <- as.vector(t(foraging$Racc_per_forager)) # aggregated resources experienced by the pollinator nesting in the focal cell
values(M_poll[[s]][[2]]) <- values(M_poll[[s]][[1]]) #Added!!!
values(M_poll[[s]][[3]]) <- values(M_poll[[s]][[2]]) * growth.func(R = values(pollres[[s]][[v]]),
a = growth[1,1,s],
b = growth[2,1,s],
max = growth[3,1,s]) #Edited!!!
values(M_poll[[s]][[1]]) <- NA #Added!!!
M_pollend = M_poll[[s]][[3]] #Added!!!
}
## ----------------- floral period 3 ----------------- ##Added!!!
v=3
##print("Floral period 3 ...");flush.console()
if(solitary_social[s]=="social" & foraging_period[s,v]=="w")
{
## N_poll (workers) aggregate resources (pollres v=2) --> VR2
## Only a proportion pw of the workers is actually foraging
foraging <- latfordisp(N = pw[s]*raster::as.matrix(N_poll[[s]][[2]]), alpha = raster::as.matrix(floral[[s]][[v]]), decay = kernelFor[[s]])
values(flowvis[[s]][[v]]) <- as.vector(t(foraging$visitation_rate)) # flower visitation in the cells surrounding the focal cell
values(pollres[[s]][[v]]) <- as.vector(t(foraging$Racc_per_forager)) #
## ------ # Population growth QUEENS # ------- ##
## M_poll produced at the end of the season depending on M_poll and resources in period 2
# growth[3,2,s] is the maximal number of workers
# N_poll is the number of foraging bees
values(M_poll[[s]][[3]]) <- values(M_poll[[s]][[1]]) * growth.func(R = values(pollres[[s]][[v]])*pw[s]*values(N_poll[[s]][[2]])/values(M_poll[[s]][[1]]), a = growth[1,1,s], b = growth[2,1,s], max = growth[3,1,s]) #Edited!!!
M_pollend = M_poll[[s]][[3]] #Added!!!
##print("growth func ran ok");flush.console()
}
if(solitary_social[s]=="solitary" & foraging_period[s,v]=="q")
{
print("Don't allow Season 3 only active solitary bees") #Added!!!
}
##print(paste("nb new females : ", round(sum(values(M_pollend))) ));flush.console() #Edited!!!
## M_poll at the beginning of the next season depending on winter survival of M_poll at the end of the season
# non-honeybees disperse to their new nesting sites #
if(wild_managed[s]=="wild")
{
poll.out.start <- latfordisp(N = raster::as.matrix(M_pollend),
alpha = raster::as.matrix(nest[[s]]),
decay = kernelNest[[s]]) #Edited!!!
values(M_poll[[s]][[(num_floral+1)]]) <- pmin(as.vector(t(poll.out.start$visitation_rate)),(av[s]*cell.size^2/10000)*values(nest[[s]])) #Edited!!!
##print(paste("nb females surv : ", round(sum(values(M_poll[[s]][[(num_floral+1)]]))) ));flush.console() #Edited!!!
}
} #end species
return(list(M_poll=M_poll,
N_poll=N_poll,
flowvis=flowvis,
floral=floral,
nest=nest,
pollres=pollres))
}
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