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## This code is part of the ips package
## Written by C. Heibl 2014 (last update 2025-09-14)
#' @title Find Pairs of Sister Species
#' @description Finds pairs of sister species in
#' a phylogenetic tree.
#' @param phy An object of class \code{\link[ape:read.tree]{phylo}}.
#' @param labels Logical, indicating whether to return tip labels or tip numbers.
#' @return A list of which each element contains the tip labels
#' of a sister species pair.
#' @examples
#' set.seed(1234)
#' tr <- rtree(12)
#' plot(tr)
#' terminalSisters(tr)
#' @export
terminalSisters <- function(phy, labels = TRUE) {
obj <- unique(
lapply(1:Ntip(phy), function(x) sort(c(x, sister(x, phy = phy))))
)
is.nested <- function(x, y){
identical(sort(union(x, y)), x)
}
id <- vapply(seq_along(obj), function(i)
!any(vapply(obj[-i], is.nested, x = obj[[i]], logical(1))),
logical(1))
obj <- obj[id]
if (labels) obj <- lapply(obj, function(phy, x) phy$tip.label[x], phy = phy)
obj
}
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