Nothing
R/cophylo.R
In phytools: Phylogenetic Tools for Comparative Biology (and Other Things)
Defines functions
summary.cophylo
drawCurve
tiplabels.cophylo
edgelabels.cophylo
nodelabels.cophylo
MULTI2DI
tipRotate
print.multiCophylo
print.cophylo
add.scalebar
plot.cophylo
plot.multiCophylo
makelinks
TEXTBOX
cladogram
phylogram
cophylo
Documented in
cophylo
edgelabels.cophylo
nodelabels.cophylo
plot.cophylo
tiplabels.cophylo
tipRotate
## creates an object of class "cophylo"
## written by Liam J. Revell 2015, 2016, 2017, 2019, 2021, 2023, 2024
cophylo<-function(tr1,tr2,assoc=NULL,rotate=TRUE,...){
if(!inherits(tr1,"phylo")||!inherits(tr2,"phylo"))
stop("tr1 & tr2 should be objects of class \"phylo\".")
## check optional arguments
if(hasArg(methods)) methods<-list(...)$methods
else methods<-"pre"
if("exhaustive"%in%methods) methods<-"exhaustive"
## hack to make sure tip labels of each tree are in cladewise order
tr1<-untangle(tr1,"read.tree")
tr2<-untangle(tr2,"read.tree")
## if no association matrix check for exact matches
if(is.null(assoc)){
assoc<-intersect(tr1$tip.label,tr2$tip.label)
assoc<-if(length(assoc)>0) cbind(assoc,assoc) else NULL
if(is.null(assoc)){
cat("No associations provided or found.\n")
rotate<-FALSE
}
}
## check to verify that all taxa in assoc are in tree
ii<-sapply(assoc[,1],"%in%",tr1$tip.label)
if(any(!ii)){
assoc<-assoc[ii,]
cat("Some species in assoc[,1] not in tr1. Removing species & links.\n")
}
ii<-sapply(assoc[,2],"%in%",tr2$tip.label)
if(any(!ii)){
assoc<-assoc[ii,]
cat("Some species in assoc[,2] not in tr2. Removing species & links.\n")
}
## now check if rotation is to be performed
if(rotate){
cat("Rotating nodes to optimize matching...\n")
flush.console()
if("exhaustive"%in%methods){
tt1<-allRotations(tr1)
tt2<-allRotations(tr2)
M1<-M2<-matrix(NA,length(tt1),length(tt2))
for(i in 1:length(tt1)){
for(j in 1:length(tt2)){
x<-setNames(sapply(assoc[,2],match,table=tt2[[j]]$tip.label),assoc[,1])
y<-setNames(sapply(assoc[,1],match,table=tt1[[i]]$tip.label),assoc[,2])
M1[i,j]<-attr(tipRotate(tt1[[i]],x*Ntip(tr1)/Ntip(tr2),methods="just.compute"),"minRotate")
M2[i,j]<-attr(tipRotate(tt2[[j]],y*Ntip(tr2)/Ntip(tr1),methods="just.compute"),"minRotate")
}
}
MM<-M1+M2
ij<-which(MM==min(MM),arr.ind=TRUE)
obj<-list()
for(i in 1:nrow(ij)){
tr1<-tt1[[ij[i,1]]]
attr(tr1,"minRotate")<-M1[ij[i,1],ij[i,2]]
tr2<-tt2[[ij[i,2]]]
attr(tr2,"minRotate")<-M2[ij[i,1],ij[i,2]]
tt<-list(tr1,tr2)
class(tt)<-"multiPhylo"
obj[[i]]<-list(trees=tt,assoc=assoc)
class(obj[[i]])<-"cophylo"
}
if(length(obj)>1) class(obj)<-"multiCophylo"
else obj<-obj[[1]]
} else if ("all"%in%methods){
tt1<-allRotations(tr1)
tt2<-allRotations(tr2)
obj<-vector(mode="list",length=length(tt1)*length(tt2))
ij<-1
for(i in 1:length(tt1)){
for(j in 1:length(tt2)){
x<-setNames(sapply(assoc[,2],match,table=tt2[[j]]$tip.label),assoc[,1])
y<-setNames(sapply(assoc[,1],match,table=tt1[[i]]$tip.label),assoc[,2])
obj[[ij]]<-list(trees=c(
tipRotate(tt1[[i]],x*Ntip(tr1)/Ntip(tr2),methods="just.compute"),
tipRotate(tt2[[j]],y*Ntip(tr2)/Ntip(tr1),methods="just.compute")),
assoc=assoc)
class(obj[[ij]])<-"cophylo"
ij<-ij+1
}
}
class(obj)<-"multiCophylo"
} else {
x<-setNames(sapply(assoc[,2],match,table=tr2$tip.label),assoc[,1])
tr1<-tipRotate(tr1,x*Ntip(tr1)/Ntip(tr2),right=tr2,assoc=assoc,...)
best.tr1<-Inf
x<-setNames(sapply(assoc[,1],match,table=tr1$tip.label),assoc[,2])
tr2<-tipRotate(tr2,x*Ntip(tr2)/Ntip(tr1),left=tr1,assoc=assoc,...)
best.tr2<-Inf
while((best.tr2-attr(tr2,"minRotate"))>0||(best.tr1-attr(tr1,"minRotate"))>0){
best.tr1<-attr(tr1,"minRotate")
best.tr2<-attr(tr2,"minRotate")
x<-setNames(sapply(assoc[,2],match,table=tr2$tip.label),assoc[,1])
tr1<-tipRotate(tr1,x*Ntip(tr1)/Ntip(tr2),right=tr2,assoc=assoc,...)
x<-setNames(sapply(assoc[,1],match,table=tr1$tip.label),assoc[,2])
tr2<-tipRotate(tr2,x*Ntip(tr2)/Ntip(tr1),left=tr1,assoc=assoc,...)
}
tt<-list(tr1,tr2)
class(tt)<-"multiPhylo"
obj<-list(trees=tt,assoc=assoc)
class(obj)<-"cophylo"
}
cat("Done.\n")
} else {
tt<-list(tr1,tr2)
class(tt)<-"multiPhylo"
obj<-list(trees=tt,assoc=assoc)
class(obj)<-"cophylo"
}
obj
}
## called internally by plot.cophylo to plot a phylogram
## written by Liam J. Revell
phylogram<-function(tree,part=1,direction="rightwards",fsize=1,ftype="i",lwd=1,...){
if(hasArg(pts)) pts<-list(...)$pts
else pts<-TRUE
if(hasArg(edge.col)) edge.col<-list(...)$edge.col
else edge.col<-rep("black",nrow(tree$edge))
if(hasArg(tip.lwd)) tip.lwd<-list(...)$tip.lwd
else tip.lwd<-1
if(hasArg(tip.lty)) tip.lty<-list(...)$tip.lty
else tip.lty<-"dotted"
if(hasArg(tip.len)) tip.len<-list(...)$tip.len
else tip.len<-0.1
if(pts==TRUE&&tip.len==0) tip.len<-0.1
if(hasArg(frame)) frame<-list(...)$frame
else frame<-TRUE
d<-if(direction=="rightwards") 1 else -1
## sub "_" for " "
tree$tip.label<-gsub("_"," ",tree$tip.label)
## check if edge lenths
if(is.null(tree$edge.length)) tree<-compute.brlen(tree)
## rescale tree so it fits in one half of the plot
## with enough space for labels
if(ftype=="off") fsize<-0
n<-Ntip(tree)
sh<-fsize*strwidth(tree$tip.label)
H<-nodeHeights(tree)
th<-sapply(1:n,function(i,x,e) x[which(e==i)],x=H[,2],
e=tree$edge[,2])+tip.len*max(H)
tree$edge.length<-tree$edge.length/max(th/(part-sh))
## reorder cladewise to assign tip positions
cw<-reorder(tree,"cladewise")
y<-vector(length=n+cw$Nnode)
y[cw$edge[cw$edge[,2]<=n,2]]<-0:(n-1)/(n-1)
## reorder pruningwise for post-order traversal
pw<-reorder(tree,"pruningwise")
nn<-unique(pw$edge[,1])
## compute vertical position of each edge
for(i in 1:length(nn)){
yy<-y[pw$edge[which(pw$edge[,1]==nn[i]),2]]
y[nn[i]]<-mean(range(yy))
}
## compute start & end points of each edge
X<-nodeHeights(cw)-0.5
## plot horizontal edges
for(i in 1:nrow(X)) lines(d*X[i,],rep(y[cw$edge[i,2]],2),lwd=lwd,lend=2,
col=edge.col[i])
## plot vertical relationships
for(i in 1:tree$Nnode+n){
ee<-which(cw$edge[,1]==i)
p<-if(i%in%cw$edge[,2]) which(cw$edge[,2]==i) else NULL
if(!is.null(p)){
xx<-c(X[ee,1],X[p,2])
yy<-sort(c(y[cw$edge[ee,2]],y[cw$edge[p,2]]))
} else {
xx<-c(X[ee,1],X[ee[1],1])
yy<-sort(c(y[cw$edge[ee,2]],mean(y[cw$edge[ee,2]])))
}
segments(x0=d*xx[1:(length(xx)-1)],y0=yy[1:(length(yy)-1)],
x1=d*xx[2:length(xx)],y1=yy[2:length(yy)],lwd=lwd,lend=2,col=edge.col[ee])
}
h<-part-0.5-tip.len*(max(X)-min(X))-fsize*strwidth(tree$tip.label)
## plot links to tips
for(i in 1:n){
lines(d*c(X[which(cw$edge[,2]==i),2],h[i]+tip.len*(max(X)-min(X))),rep(y[i],2),
lwd=tip.lwd,lty=tip.lty)
if(pts) points(d*X[which(cw$edge[,2]==i),2],y[i],pch=16,cex=pts*0.7*sqrt(lwd))
}
## plot tip labels
font<-which(c("off","reg","b","i","bi")==ftype)-1
if(font>0){
for(i in 1:n) TEXTBOX(d*(h[i]+fsize*strwidth(tree$tip.label[i])+
tip.len*(max(X)-min(X))),y[i],
tree$tip.label[i], pos=if(d<0) 4 else 2,offset=0,
cex=fsize,font=font,frame=frame)
}
PP<-list(type="phylogram",use.edge.length=TRUE,node.pos=1,
show.tip.label=if(ftype!="off") TRUE else FALSE,show.node.label=FALSE,
font=ftype,cex=fsize,adj=0,srt=0,no.margin=FALSE,label.offset=0,
x.lim=par()$usr[1:2],y.lim=par()$usr[3:4],
direction=direction,tip.color="black",Ntip=Ntip(cw),Nnode=cw$Nnode,
edge=tree$edge,xx=d*sapply(1:(Ntip(cw)+cw$Nnode),
function(x,y,z) y[match(x,z)],y=X,z=cw$edge),yy=y)
assign("last_plot.phylo",PP,envir=.PlotPhyloEnv)
## return rightmost or leftmost edge of tip labels
invisible(d*max(h+fsize*strwidth(tree$tip.label)+tip.len*(max(X)-min(X))))
}
cladogram<-function(tree,part=1,direction="rightwards",fsize=1,ftype="i",lwd=1,...){
if(hasArg(pts)) pts<-list(...)$pts
else pts<-TRUE
if(hasArg(edge.col)) edge.col<-list(...)$edge.col
else edge.col<-rep("black",nrow(tree$edge))
if(hasArg(tip.lwd)) tip.lwd<-list(...)$tip.lwd
else tip.lwd<-1
if(hasArg(tip.lty)) tip.lty<-list(...)$tip.lty
else tip.lty<-"dotted"
if(hasArg(tip.len)) tip.len<-list(...)$tip.len
else tip.len<-0.1
if(pts==TRUE&&tip.len==0) tip.len<-0.1
if(hasArg(frame)) frame<-list(...)$frame
else frame<-TRUE
d<-if(direction=="rightwards") 1 else -1
## sub "_" for " "
tree$tip.label<-gsub("_"," ",tree$tip.label)
## check if edge lenths
if(is.null(tree$edge.length)) tree<-compute.brlen(tree)
## rescale tree so it fits in one half of the plot
## with enough space for labels
if(ftype=="off") fsize<-0
n<-Ntip(tree)
sh<-fsize*strwidth(tree$tip.label)
H<-nodeHeights(tree)
th<-sapply(1:n,function(i,x,e) x[which(e==i)],x=H[,2],
e=tree$edge[,2])+tip.len*max(H)
tree$edge.length<-tree$edge.length/max(th/(part-sh))
## reorder cladewise to assign tip positions
cw<-reorder(tree,"cladewise")
y<-vector(length=n+cw$Nnode)
y[cw$edge[cw$edge[,2]<=n,2]]<-0:(n-1)/(n-1)
## reorder pruningwise for post-order traversal
pw<-reorder(tree,"pruningwise")
nn<-unique(pw$edge[,1])
## compute vertical position of each edge
for(i in 1:length(nn)){
desc<-pw$edge[which(pw$edge[,1]==nn[i]),2]
n1<-desc[which(y[desc]==min(y[desc]))]
n2<-desc[which(y[desc]==max(y[desc]))]
v1<-pw$edge.length[which(pw$edge[,2]==n1)]
v2<-pw$edge.length[which(pw$edge[,2]==n2)]
y[nn[i]]<-((1/v1)*y[n1]+(1/v2)*y[n2])/(1/v1+1/v2)
}
## compute start & end points of each edge
X<-nodeHeights(cw)-0.5
## plot horizontal edges
for(i in 1:nrow(X)) lines(d*X[i,],y[cw$edge[i,]],lwd=lwd,lend=2,
col=edge.col[i])
h<-part-0.5-tip.len*(max(X)-min(X))-fsize*strwidth(tree$tip.label)
## plot links to tips
for(i in 1:n){
lines(d*c(X[which(cw$edge[,2]==i),2],h[i]+tip.len*(max(X)-min(X))),rep(y[i],2),
lwd=tip.lwd,lty=tip.lty)
if(pts) points(d*X[which(cw$edge[,2]==i),2],y[i],pch=16,cex=pts*0.7*sqrt(lwd))
}
## plot tip labels
font<-which(c("off","reg","b","i","bi")==ftype)-1
if(font>0){
for(i in 1:n) TEXTBOX(d*(h[i]+fsize*strwidth(tree$tip.label[i])+
tip.len*(max(X)-min(X))),y[i],
tree$tip.label[i], pos=if(d<0) 4 else 2,offset=0,
cex=fsize,font=font,frame=frame)
}
PP<-list(type="cladogram",use.edge.length=TRUE,node.pos=1,
show.tip.label=if(ftype!="off") TRUE else FALSE,show.node.label=FALSE,
font=ftype,cex=fsize,adj=0,srt=0,no.margin=FALSE,label.offset=0,
x.lim=par()$usr[1:2],y.lim=par()$usr[3:4],
direction=direction,tip.color="black",Ntip=Ntip(cw),Nnode=cw$Nnode,
edge=tree$edge,xx=d*sapply(1:(Ntip(cw)+cw$Nnode),
function(x,y,z) y[match(x,z)],y=X,z=cw$edge),yy=y)
assign("last_plot.phylo",PP,envir=.PlotPhyloEnv)
## return rightmost or leftmost edge of tip labels
invisible(d*max(h+fsize*strwidth(tree$tip.label)+tip.len*(max(X)-min(X))))
}
## internally used function
TEXTBOX<-function(x,y,label,pos,offset,cex,font,frame=TRUE){
if(frame) rect(x,y-0.5*strheight(label,cex=cex,font=font),
x+if(pos==4) strwidth(label,
cex=cex,font=font) else -strwidth(label,cex=cex,font=font),
y+0.5*strheight(label,cex=cex,font=font),border=FALSE,
col=if(par()$bg%in%c("white","transparent")) "white" else par()$bg)
text(x=x,y=y,label=label,pos=pos,offset=offset,cex=cex,font=font)
}
## plot links between tip taxa according to assoc
## written by Liam J. Revell 2015, 2016, 2019
makelinks<-function(obj,x,link.type="curved",link.lwd=1,link.col="black",
link.lty="dashed"){
if(length(link.lwd)==1) link.lwd<-rep(link.lwd,nrow(obj$assoc))
if(length(link.col)==1) link.col<-rep(link.col,nrow(obj$assoc))
if(length(link.lty)==1) link.lty<-rep(link.lty,nrow(obj$assoc))
for(i in 1:nrow(obj$assoc)){
ii<-which(obj$trees[[1]]$tip.label==obj$assoc[i,1])
jj<-which(obj$trees[[2]]$tip.label==obj$assoc[i,2])
for(j in 1:length(ii)) for(k in 1:length(jj)){
y<-c((ii[j]-1)/(Ntip(obj$trees[[1]])-1),
(jj[k]-1)/(Ntip(obj$trees[[2]])-1))
if(link.type=="straight") lines(x,y,lty=link.lty[i],
lwd=link.lwd[i],col=link.col[i])
else if(link.type=="curved") drawCurve(x,y,lty=link.lty[i],
lwd=link.lwd[i],col=link.col[i])
}
}
}
## plot method for class "multiCophylo"
plot.multiCophylo<-function(x,...){
par(ask=TRUE)
for(i in 1:length(x)) plot.cophylo(x[[i]],...)
}
## plot an object of class "cophylo"
## written by Liam J. Revell 2015, 2016, 2017, 2021, 2023
plot.cophylo<-function(x,...){
plot.new()
if(hasArg(type)) type<-list(...)$type
else type<-"phylogram"
if(length(type)==1) type<-rep(type,2)
if(hasArg(mar)) mar<-list(...)$mar
else mar<-c(0.1,0.1,0.1,0.1)
if(hasArg(xlim)) xlim<-list(...)$xlim
else xlim<-c(-0.5,0.5)
if(hasArg(scale.bar)) scale.bar<-list(...)$scale.bar
else scale.bar<-rep(0,2)
if(hasArg(ylim)) ylim<-list(...)$ylim
else ylim<-if(any(scale.bar>0)) c(-0.1,1) else c(0,1)
if(hasArg(link.type)) link.type<-list(...)$link.type
else link.type<-"straight"
if(hasArg(link.lwd)) link.lwd<-list(...)$link.lwd
else link.lwd<-1
if(hasArg(link.col)) link.col<-list(...)$link.col
else link.col<-"black"
if(hasArg(link.lty)) link.lty<-list(...)$link.lty
else link.lty<-"dashed"
if(hasArg(edge.col)) edge.col<-list(...)$edge.col
else edge.col<-list(
left=rep("black",nrow(x$trees[[1]]$edge)),
right=rep("black",nrow(x$trees[[2]]$edge)))
if(hasArg(frame)) frame<-list(...)$frame
else frame<-c(TRUE,TRUE)
if(length(frame)==1) frame<-rep(frame,2)
obj<-list(...)
if(is.null(obj$part)) obj$part<-0.4
par(mar=mar)
plot.window(xlim=xlim,ylim=ylim)
leftArgs<-rightArgs<-obj
leftArgs$edge.col<-edge.col$left
rightArgs$edge.col<-edge.col$right
if(!is.null(obj$fsize)){
if(length(obj$fsize)>1){
leftArgs$fsize<-obj$fsize[1]
rightArgs$fsize<-obj$fsize[2]
sb.fsize<- if(length(obj$fsize)>2) obj$fsize[3] else 1
} else sb.fsize<-1
} else sb.fsize<-1
if(!is.null(obj$ftype)){
if(length(obj$ftype)>1){
leftArgs$ftype<-obj$ftype[1]
rightArgs$ftype<-obj$ftype[2]
}
}
leftArgs$frame<-frame[1]
rightArgs$frame<-frame[2]
plotter<-if(type[1]=="cladogram") "cladogram" else "phylogram"
x1<-do.call(plotter,c(list(tree=x$trees[[1]]),leftArgs))
plotter<-if(type[2]=="cladogram") "cladogram" else "phylogram"
left<-get("last_plot.phylo",envir=.PlotPhyloEnv)
x2<-do.call(plotter,c(list(tree=x$trees[[2]],direction="leftwards"),
rightArgs))
right<-get("last_plot.phylo",envir=.PlotPhyloEnv)
if(!is.null(x$assoc)) makelinks(x,c(x1,x2),link.type,link.lwd,link.col,
link.lty)
else cat("No associations provided.\n")
assign("last_plot.cophylo",list(left=left,right=right),envir=.PlotPhyloEnv)
if(any(scale.bar>0)) add.scalebar(x,scale.bar,sb.fsize)
}
## add scale bar
## written by Liam J. Revell 2015, 2023
add.scalebar<-function(obj,scale.bar,fsize){
if(scale.bar[1]>0){
pp<-get("last_plot.cophylo",envir=.PlotPhyloEnv)[[1]]
s1<-diff(range(pp$xx))/max(nodeHeights(obj$trees[[1]]))
lines(c(-0.5,-0.5+scale.bar[1]*s1),rep(-0.05,2))
lines(rep(-0.5,2),c(-0.05,-0.06))
lines(rep(-0.5+scale.bar[1]*s1,2),c(-0.05,-0.06))
text(mean(c(-0.5,-0.5+scale.bar[1]*s1)),rep(-0.05,2),scale.bar[1],pos=1)
}
if(scale.bar[2]>0){
pp<-get("last_plot.cophylo",envir=.PlotPhyloEnv)[[2]]
s2<-diff(range(pp$xx))/max(nodeHeights(obj$trees[[2]]))
lines(c(0.5-scale.bar[2]*s2,0.5),rep(-0.05,2))
lines(rep(0.5-scale.bar[2]*s2,2),c(-0.05,-0.06))
lines(rep(0.5,2),c(-0.05,-0.06))
text(mean(c(0.5-scale.bar[2]*s2,0.5)),rep(-0.05,2),scale.bar[2],pos=1)
}
}
## print an object of class "cophylo"
## written by Liam J. Revell 2015
print.cophylo<-function(x, ...){
cat("Object of class \"cophylo\" containing:\n\n")
cat("(1) 2 (possibly rotated) phylogenetic trees in an object of class \"multiPhylo\".\n\n")
cat("(2) A table of associations between the tips of both trees.\n\n")
}
## print method for "multiCophylo" object
print.multiCophylo<-function(x, ...)
cat("Object of class \"multiCophylo\" containg",length(x),"objects of class \"cophylo\".\n\n")
## written by Liam J. Revell 2015, 2016
tipRotate<-function(tree,x,...){
if(hasArg(fn)) fn<-list(...)$fn
else fn<-function(x) x^2
if(hasArg(methods)) methods<-list(...)$methods
else methods<-"pre"
if("exhaustive"%in%methods) methods<-"exhaustive"
if(hasArg(print)) print<-list(...)$print
else print<-FALSE
if(hasArg(max.exhaustive)) max.exhaustive<-list(...)$max.exhaustive
else max.exhaustive<-20
if(hasArg(rotate.multi)) rotate.multi<-list(...)$rotate.multi
else rotate.multi<-FALSE
if(rotate.multi) rotate.multi<-!is.binary(tree)
if(hasArg(anim.cophylo)) anim.cophylo<-list(...)$anim.cophylo
else anim.cophylo<-FALSE
if(anim.cophylo){
if(hasArg(left)) left<-list(...)$left
else left<-NULL
if(hasArg(right)) right<-list(...)$right
else right<-NULL
if(hasArg(assoc)) assoc<-list(...)$assoc
else assoc<-NULL
if(is.null(left)&&is.null(right)) anim.cophylo<-FALSE
if(hasArg(only.accepted)) only.accepted<-list(...)$only.accepted
else only.accepted<-TRUE
}
tree<-reorder(tree)
nn<-1:tree$Nnode+length(tree$tip.label)
if("just.compute"%in%methods){
foo<-function(phy,x) sum(fn(x-setNames(1:length(phy$tip.label),phy$tip.label)[names(x)]))
oo<-pp<-foo(tree,x)
}
if("exhaustive"%in%methods){
if(Ntip(tree)>max.exhaustive){
cat(paste("\nmethods=\"exhaustive\" not permitted for more than",
max.exhaustive,"tips.\n",
"If you are sure you want to run an exhaustive search for a tree of this size\n",
"increasing argument max.exhaustive & re-run.\n",
"Setting methods to \"pre\".\n\n"))
methods<-"pre"
} else {
cat("Running exhaustive search. May be slow!\n")
oo<-Inf
tt<-allRotations(tree)
foo<-function(phy,x) sum(fn(x-setNames(1:length(phy$tip.label),phy$tip.label)[names(x)]))
pp<-sapply(tt,foo,x=x)
ii<-which(pp==min(pp))
ii<-if(length(ii)>1) sample(ii,1) else ii
tt<-tt[[ii]]
pp<-pp[ii]
}
if(print) message(paste("objective:",pp))
tree<-tt
}
ANIM.COPHYLO<-function(tree){
dev.hold()
if(is.null(left)) plot(cophylo(tree,right,assoc=assoc,rotate=FALSE),...)
else if(is.null(right)) plot(cophylo(left,tree,assoc=assoc,rotate=FALSE),...)
nodelabels.cophylo(node=i+Ntip(tree),pie=1,col="red",cex=0.4,
which=if(is.null(left)) "left" else "right")
dev.flush()
}
if("pre"%in%methods){
for(i in 1:tree$Nnode){
if(anim.cophylo) ANIM.COPHYLO(tree)
tt<-if(rotate.multi) rotate.multi(tree,nn[i]) else untangle(rotate(tree,nn[i]),"read.tree")
oo<-sum(fn(x-setNames(1:length(tree$tip.label),tree$tip.label)[names(x)]))
if(inherits(tt,"phylo")) pp<-sum(fn(x-setNames(1:length(tt$tip.label),tt$tip.label)[names(x)]))
if(anim.cophylo&&!only.accepted) ANIM.COPHYLO(tt)
else if(inherits(tt,"multiPhylo")){
foo<-function(phy,x) sum(fn(x-setNames(1:length(phy$tip.label),phy$tip.label)[names(x)]))
pp<-sapply(tt,foo,x=x)
if(anim.cophylo&&!only.accepted) nulo<-lapply(tt,ANIM.COPHYLO)
ii<-which(pp==min(pp))
ii<-if(length(ii)>1) sample(ii,1) else ii
tt<-tt[[ii]]
pp<-pp[ii]
}
if(oo>pp) tree<-tt
if(print) message(paste("objective:",min(oo,pp)))
}
}
if("post"%in%methods){
for(i in tree$Nnode:1){
if(anim.cophylo) ANIM.COPHYLO(tree)
tt<-if(rotate.multi) rotate.multi(tree,nn[i]) else untangle(rotate(tree,nn[i]),"read.tree")
oo<-sum(fn(x-setNames(1:length(tree$tip.label),tree$tip.label)[names(x)]))
if(inherits(tt,"phylo")) pp<-sum(fn(x-setNames(1:length(tt$tip.label),tt$tip.label)[names(x)]))
if(anim.cophylo&&!only.accepted) ANIM.COPHYLO(tt)
else if(inherits(tt,"multiPhylo")){
foo<-function(phy,x) sum(fn(x-setNames(1:length(phy$tip.label),phy$tip.label)[names(x)]))
pp<-sapply(tt,foo,x=x)
if(anim.cophylo&&!only.accepted) nulo<-lapply(tt,ANIM.COPHYLO)
ii<-which(pp==min(pp))
ii<-if(length(ii)>1) sample(ii,1) else ii
tt<-tt[[ii]]
pp<-pp[ii]
}
if(oo>pp) tree<-tt
if(print) message(paste("objective:",min(oo,pp)))
}
}
attr(tree,"minRotate")<-min(oo,pp)
if(anim.cophylo) ANIM.COPHYLO(tree)
tree
}
## multi2di for "multiPhylo" object
MULTI2DI<-function(x,...){
obj<-lapply(x,multi2di,...)
class(obj)<-"multiPhylo"
obj
}
## labeling methods for plotted "cophylo" object
## written by Liam J. Revell 2015
nodelabels.cophylo<-function(...,which=c("left","right")){
obj<-get("last_plot.cophylo",envir=.PlotPhyloEnv)
if(which[1]=="left") assign("last_plot.phylo",obj[[1]],envir=.PlotPhyloEnv)
else if(which[1]=="right") assign("last_plot.phylo",obj[[2]],envir=.PlotPhyloEnv)
nodelabels(...)
}
edgelabels.cophylo<-function(...,which=c("left","right")){
obj<-get("last_plot.cophylo",envir=.PlotPhyloEnv)
if(which[1]=="left") assign("last_plot.phylo",obj[[1]],envir=.PlotPhyloEnv)
else if(which[1]=="right") assign("last_plot.phylo",obj[[2]],envir=.PlotPhyloEnv)
edgelabels(...)
}
tiplabels.cophylo<-function(...,which=c("left","right")){
obj<-get("last_plot.cophylo",envir=.PlotPhyloEnv)
if(which[1]=="left") assign("last_plot.phylo",obj[[1]],envir=.PlotPhyloEnv)
else if(which[1]=="right") assign("last_plot.phylo",obj[[2]],envir=.PlotPhyloEnv)
tiplabels(...)
}
## function to draw sigmoidal links
## modified from https://stackoverflow.com/questions/32046889/connecting-two-points-with-curved-lines-s-ish-curve-in-r
drawCurve<-function(x,y,scale=0.01,...){
x1<-x[1]
x2<-x[2]
y1<-y[1]
y2<-y[2]
curve(plogis(x,scale=scale,location=(x1+x2)/2)*(y2-y1)+y1,
x1,x2,add=TRUE,...)
}
## S3 summary method
## written by Liam J. Revell 2016
summary.cophylo<-function(object,...){
cat("\nCo-phylogenetic (\"cophylo\") object:",deparse(substitute(object)),
"\n\n")
cat(paste("Tree 1 (left tree) is an object of class \"phylo\" containing",
Ntip(object$trees[[1]]),"species.\n\n"))
cat(paste("Tree 2 (right tree) is an object of class \"phylo\" containing",
Ntip(object$trees[[2]]),"species.\n\n"))
cat("Association (assoc) table as follows:\n\n")
maxl<-max(sapply(strsplit(object$assoc[,1],""),length))
cat(paste("\tleft:",paste(rep(" ",max(0,maxl-5)),collapse=""),
"\t----\tright:\n",sep=""))
nulo<-apply(object$assoc,1,function(x,maxl) cat(paste("\t",x[1],
paste(rep(" ",maxl-length(strsplit(x[1],split="")[[1]])),
collapse=""),"\t----\t",x[2],"\n",sep="")),maxl=maxl)
cat("\n")
}
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Defines functions summary.cophylo drawCurve tiplabels.cophylo edgelabels.cophylo nodelabels.cophylo MULTI2DI tipRotate print.multiCophylo print.cophylo add.scalebar plot.cophylo plot.multiCophylo makelinks TEXTBOX cladogram phylogram cophylo
Documented in cophylo edgelabels.cophylo nodelabels.cophylo plot.cophylo tiplabels.cophylo tipRotate
## creates an object of class "cophylo"
## written by Liam J. Revell 2015, 2016, 2017, 2019, 2021, 2023, 2024
cophylo<-function(tr1,tr2,assoc=NULL,rotate=TRUE,...){
if(!inherits(tr1,"phylo")||!inherits(tr2,"phylo"))
stop("tr1 & tr2 should be objects of class \"phylo\".")
## check optional arguments
if(hasArg(methods)) methods<-list(...)$methods
else methods<-"pre"
if("exhaustive"%in%methods) methods<-"exhaustive"
## hack to make sure tip labels of each tree are in cladewise order
tr1<-untangle(tr1,"read.tree")
tr2<-untangle(tr2,"read.tree")
## if no association matrix check for exact matches
if(is.null(assoc)){
assoc<-intersect(tr1$tip.label,tr2$tip.label)
assoc<-if(length(assoc)>0) cbind(assoc,assoc) else NULL
if(is.null(assoc)){
cat("No associations provided or found.\n")
rotate<-FALSE
}
}
## check to verify that all taxa in assoc are in tree
ii<-sapply(assoc[,1],"%in%",tr1$tip.label)
if(any(!ii)){
assoc<-assoc[ii,]
cat("Some species in assoc[,1] not in tr1. Removing species & links.\n")
}
ii<-sapply(assoc[,2],"%in%",tr2$tip.label)
if(any(!ii)){
assoc<-assoc[ii,]
cat("Some species in assoc[,2] not in tr2. Removing species & links.\n")
}
## now check if rotation is to be performed
if(rotate){
cat("Rotating nodes to optimize matching...\n")
flush.console()
if("exhaustive"%in%methods){
tt1<-allRotations(tr1)
tt2<-allRotations(tr2)
M1<-M2<-matrix(NA,length(tt1),length(tt2))
for(i in 1:length(tt1)){
for(j in 1:length(tt2)){
x<-setNames(sapply(assoc[,2],match,table=tt2[[j]]$tip.label),assoc[,1])
y<-setNames(sapply(assoc[,1],match,table=tt1[[i]]$tip.label),assoc[,2])
M1[i,j]<-attr(tipRotate(tt1[[i]],x*Ntip(tr1)/Ntip(tr2),methods="just.compute"),"minRotate")
M2[i,j]<-attr(tipRotate(tt2[[j]],y*Ntip(tr2)/Ntip(tr1),methods="just.compute"),"minRotate")
}
}
MM<-M1+M2
ij<-which(MM==min(MM),arr.ind=TRUE)
obj<-list()
for(i in 1:nrow(ij)){
tr1<-tt1[[ij[i,1]]]
attr(tr1,"minRotate")<-M1[ij[i,1],ij[i,2]]
tr2<-tt2[[ij[i,2]]]
attr(tr2,"minRotate")<-M2[ij[i,1],ij[i,2]]
tt<-list(tr1,tr2)
class(tt)<-"multiPhylo"
obj[[i]]<-list(trees=tt,assoc=assoc)
class(obj[[i]])<-"cophylo"
}
if(length(obj)>1) class(obj)<-"multiCophylo"
else obj<-obj[[1]]
} else if ("all"%in%methods){
tt1<-allRotations(tr1)
tt2<-allRotations(tr2)
obj<-vector(mode="list",length=length(tt1)*length(tt2))
ij<-1
for(i in 1:length(tt1)){
for(j in 1:length(tt2)){
x<-setNames(sapply(assoc[,2],match,table=tt2[[j]]$tip.label),assoc[,1])
y<-setNames(sapply(assoc[,1],match,table=tt1[[i]]$tip.label),assoc[,2])
obj[[ij]]<-list(trees=c(
tipRotate(tt1[[i]],x*Ntip(tr1)/Ntip(tr2),methods="just.compute"),
tipRotate(tt2[[j]],y*Ntip(tr2)/Ntip(tr1),methods="just.compute")),
assoc=assoc)
class(obj[[ij]])<-"cophylo"
ij<-ij+1
}
}
class(obj)<-"multiCophylo"
} else {
x<-setNames(sapply(assoc[,2],match,table=tr2$tip.label),assoc[,1])
tr1<-tipRotate(tr1,x*Ntip(tr1)/Ntip(tr2),right=tr2,assoc=assoc,...)
best.tr1<-Inf
x<-setNames(sapply(assoc[,1],match,table=tr1$tip.label),assoc[,2])
tr2<-tipRotate(tr2,x*Ntip(tr2)/Ntip(tr1),left=tr1,assoc=assoc,...)
best.tr2<-Inf
while((best.tr2-attr(tr2,"minRotate"))>0||(best.tr1-attr(tr1,"minRotate"))>0){
best.tr1<-attr(tr1,"minRotate")
best.tr2<-attr(tr2,"minRotate")
x<-setNames(sapply(assoc[,2],match,table=tr2$tip.label),assoc[,1])
tr1<-tipRotate(tr1,x*Ntip(tr1)/Ntip(tr2),right=tr2,assoc=assoc,...)
x<-setNames(sapply(assoc[,1],match,table=tr1$tip.label),assoc[,2])
tr2<-tipRotate(tr2,x*Ntip(tr2)/Ntip(tr1),left=tr1,assoc=assoc,...)
}
tt<-list(tr1,tr2)
class(tt)<-"multiPhylo"
obj<-list(trees=tt,assoc=assoc)
class(obj)<-"cophylo"
}
cat("Done.\n")
} else {
tt<-list(tr1,tr2)
class(tt)<-"multiPhylo"
obj<-list(trees=tt,assoc=assoc)
class(obj)<-"cophylo"
}
obj
}
## called internally by plot.cophylo to plot a phylogram
## written by Liam J. Revell
phylogram<-function(tree,part=1,direction="rightwards",fsize=1,ftype="i",lwd=1,...){
if(hasArg(pts)) pts<-list(...)$pts
else pts<-TRUE
if(hasArg(edge.col)) edge.col<-list(...)$edge.col
else edge.col<-rep("black",nrow(tree$edge))
if(hasArg(tip.lwd)) tip.lwd<-list(...)$tip.lwd
else tip.lwd<-1
if(hasArg(tip.lty)) tip.lty<-list(...)$tip.lty
else tip.lty<-"dotted"
if(hasArg(tip.len)) tip.len<-list(...)$tip.len
else tip.len<-0.1
if(pts==TRUE&&tip.len==0) tip.len<-0.1
if(hasArg(frame)) frame<-list(...)$frame
else frame<-TRUE
d<-if(direction=="rightwards") 1 else -1
## sub "_" for " "
tree$tip.label<-gsub("_"," ",tree$tip.label)
## check if edge lenths
if(is.null(tree$edge.length)) tree<-compute.brlen(tree)
## rescale tree so it fits in one half of the plot
## with enough space for labels
if(ftype=="off") fsize<-0
n<-Ntip(tree)
sh<-fsize*strwidth(tree$tip.label)
H<-nodeHeights(tree)
th<-sapply(1:n,function(i,x,e) x[which(e==i)],x=H[,2],
e=tree$edge[,2])+tip.len*max(H)
tree$edge.length<-tree$edge.length/max(th/(part-sh))
## reorder cladewise to assign tip positions
cw<-reorder(tree,"cladewise")
y<-vector(length=n+cw$Nnode)
y[cw$edge[cw$edge[,2]<=n,2]]<-0:(n-1)/(n-1)
## reorder pruningwise for post-order traversal
pw<-reorder(tree,"pruningwise")
nn<-unique(pw$edge[,1])
## compute vertical position of each edge
for(i in 1:length(nn)){
yy<-y[pw$edge[which(pw$edge[,1]==nn[i]),2]]
y[nn[i]]<-mean(range(yy))
}
## compute start & end points of each edge
X<-nodeHeights(cw)-0.5
## plot horizontal edges
for(i in 1:nrow(X)) lines(d*X[i,],rep(y[cw$edge[i,2]],2),lwd=lwd,lend=2,
col=edge.col[i])
## plot vertical relationships
for(i in 1:tree$Nnode+n){
ee<-which(cw$edge[,1]==i)
p<-if(i%in%cw$edge[,2]) which(cw$edge[,2]==i) else NULL
if(!is.null(p)){
xx<-c(X[ee,1],X[p,2])
yy<-sort(c(y[cw$edge[ee,2]],y[cw$edge[p,2]]))
} else {
xx<-c(X[ee,1],X[ee[1],1])
yy<-sort(c(y[cw$edge[ee,2]],mean(y[cw$edge[ee,2]])))
}
segments(x0=d*xx[1:(length(xx)-1)],y0=yy[1:(length(yy)-1)],
x1=d*xx[2:length(xx)],y1=yy[2:length(yy)],lwd=lwd,lend=2,col=edge.col[ee])
}
h<-part-0.5-tip.len*(max(X)-min(X))-fsize*strwidth(tree$tip.label)
## plot links to tips
for(i in 1:n){
lines(d*c(X[which(cw$edge[,2]==i),2],h[i]+tip.len*(max(X)-min(X))),rep(y[i],2),
lwd=tip.lwd,lty=tip.lty)
if(pts) points(d*X[which(cw$edge[,2]==i),2],y[i],pch=16,cex=pts*0.7*sqrt(lwd))
}
## plot tip labels
font<-which(c("off","reg","b","i","bi")==ftype)-1
if(font>0){
for(i in 1:n) TEXTBOX(d*(h[i]+fsize*strwidth(tree$tip.label[i])+
tip.len*(max(X)-min(X))),y[i],
tree$tip.label[i], pos=if(d<0) 4 else 2,offset=0,
cex=fsize,font=font,frame=frame)
}
PP<-list(type="phylogram",use.edge.length=TRUE,node.pos=1,
show.tip.label=if(ftype!="off") TRUE else FALSE,show.node.label=FALSE,
font=ftype,cex=fsize,adj=0,srt=0,no.margin=FALSE,label.offset=0,
x.lim=par()$usr[1:2],y.lim=par()$usr[3:4],
direction=direction,tip.color="black",Ntip=Ntip(cw),Nnode=cw$Nnode,
edge=tree$edge,xx=d*sapply(1:(Ntip(cw)+cw$Nnode),
function(x,y,z) y[match(x,z)],y=X,z=cw$edge),yy=y)
assign("last_plot.phylo",PP,envir=.PlotPhyloEnv)
## return rightmost or leftmost edge of tip labels
invisible(d*max(h+fsize*strwidth(tree$tip.label)+tip.len*(max(X)-min(X))))
}
cladogram<-function(tree,part=1,direction="rightwards",fsize=1,ftype="i",lwd=1,...){
if(hasArg(pts)) pts<-list(...)$pts
else pts<-TRUE
if(hasArg(edge.col)) edge.col<-list(...)$edge.col
else edge.col<-rep("black",nrow(tree$edge))
if(hasArg(tip.lwd)) tip.lwd<-list(...)$tip.lwd
else tip.lwd<-1
if(hasArg(tip.lty)) tip.lty<-list(...)$tip.lty
else tip.lty<-"dotted"
if(hasArg(tip.len)) tip.len<-list(...)$tip.len
else tip.len<-0.1
if(pts==TRUE&&tip.len==0) tip.len<-0.1
if(hasArg(frame)) frame<-list(...)$frame
else frame<-TRUE
d<-if(direction=="rightwards") 1 else -1
## sub "_" for " "
tree$tip.label<-gsub("_"," ",tree$tip.label)
## check if edge lenths
if(is.null(tree$edge.length)) tree<-compute.brlen(tree)
## rescale tree so it fits in one half of the plot
## with enough space for labels
if(ftype=="off") fsize<-0
n<-Ntip(tree)
sh<-fsize*strwidth(tree$tip.label)
H<-nodeHeights(tree)
th<-sapply(1:n,function(i,x,e) x[which(e==i)],x=H[,2],
e=tree$edge[,2])+tip.len*max(H)
tree$edge.length<-tree$edge.length/max(th/(part-sh))
## reorder cladewise to assign tip positions
cw<-reorder(tree,"cladewise")
y<-vector(length=n+cw$Nnode)
y[cw$edge[cw$edge[,2]<=n,2]]<-0:(n-1)/(n-1)
## reorder pruningwise for post-order traversal
pw<-reorder(tree,"pruningwise")
nn<-unique(pw$edge[,1])
## compute vertical position of each edge
for(i in 1:length(nn)){
desc<-pw$edge[which(pw$edge[,1]==nn[i]),2]
n1<-desc[which(y[desc]==min(y[desc]))]
n2<-desc[which(y[desc]==max(y[desc]))]
v1<-pw$edge.length[which(pw$edge[,2]==n1)]
v2<-pw$edge.length[which(pw$edge[,2]==n2)]
y[nn[i]]<-((1/v1)*y[n1]+(1/v2)*y[n2])/(1/v1+1/v2)
}
## compute start & end points of each edge
X<-nodeHeights(cw)-0.5
## plot horizontal edges
for(i in 1:nrow(X)) lines(d*X[i,],y[cw$edge[i,]],lwd=lwd,lend=2,
col=edge.col[i])
h<-part-0.5-tip.len*(max(X)-min(X))-fsize*strwidth(tree$tip.label)
## plot links to tips
for(i in 1:n){
lines(d*c(X[which(cw$edge[,2]==i),2],h[i]+tip.len*(max(X)-min(X))),rep(y[i],2),
lwd=tip.lwd,lty=tip.lty)
if(pts) points(d*X[which(cw$edge[,2]==i),2],y[i],pch=16,cex=pts*0.7*sqrt(lwd))
}
## plot tip labels
font<-which(c("off","reg","b","i","bi")==ftype)-1
if(font>0){
for(i in 1:n) TEXTBOX(d*(h[i]+fsize*strwidth(tree$tip.label[i])+
tip.len*(max(X)-min(X))),y[i],
tree$tip.label[i], pos=if(d<0) 4 else 2,offset=0,
cex=fsize,font=font,frame=frame)
}
PP<-list(type="cladogram",use.edge.length=TRUE,node.pos=1,
show.tip.label=if(ftype!="off") TRUE else FALSE,show.node.label=FALSE,
font=ftype,cex=fsize,adj=0,srt=0,no.margin=FALSE,label.offset=0,
x.lim=par()$usr[1:2],y.lim=par()$usr[3:4],
direction=direction,tip.color="black",Ntip=Ntip(cw),Nnode=cw$Nnode,
edge=tree$edge,xx=d*sapply(1:(Ntip(cw)+cw$Nnode),
function(x,y,z) y[match(x,z)],y=X,z=cw$edge),yy=y)
assign("last_plot.phylo",PP,envir=.PlotPhyloEnv)
## return rightmost or leftmost edge of tip labels
invisible(d*max(h+fsize*strwidth(tree$tip.label)+tip.len*(max(X)-min(X))))
}
## internally used function
TEXTBOX<-function(x,y,label,pos,offset,cex,font,frame=TRUE){
if(frame) rect(x,y-0.5*strheight(label,cex=cex,font=font),
x+if(pos==4) strwidth(label,
cex=cex,font=font) else -strwidth(label,cex=cex,font=font),
y+0.5*strheight(label,cex=cex,font=font),border=FALSE,
col=if(par()$bg%in%c("white","transparent")) "white" else par()$bg)
text(x=x,y=y,label=label,pos=pos,offset=offset,cex=cex,font=font)
}
## plot links between tip taxa according to assoc
## written by Liam J. Revell 2015, 2016, 2019
makelinks<-function(obj,x,link.type="curved",link.lwd=1,link.col="black",
link.lty="dashed"){
if(length(link.lwd)==1) link.lwd<-rep(link.lwd,nrow(obj$assoc))
if(length(link.col)==1) link.col<-rep(link.col,nrow(obj$assoc))
if(length(link.lty)==1) link.lty<-rep(link.lty,nrow(obj$assoc))
for(i in 1:nrow(obj$assoc)){
ii<-which(obj$trees[[1]]$tip.label==obj$assoc[i,1])
jj<-which(obj$trees[[2]]$tip.label==obj$assoc[i,2])
for(j in 1:length(ii)) for(k in 1:length(jj)){
y<-c((ii[j]-1)/(Ntip(obj$trees[[1]])-1),
(jj[k]-1)/(Ntip(obj$trees[[2]])-1))
if(link.type=="straight") lines(x,y,lty=link.lty[i],
lwd=link.lwd[i],col=link.col[i])
else if(link.type=="curved") drawCurve(x,y,lty=link.lty[i],
lwd=link.lwd[i],col=link.col[i])
}
}
}
## plot method for class "multiCophylo"
plot.multiCophylo<-function(x,...){
par(ask=TRUE)
for(i in 1:length(x)) plot.cophylo(x[[i]],...)
}
## plot an object of class "cophylo"
## written by Liam J. Revell 2015, 2016, 2017, 2021, 2023
plot.cophylo<-function(x,...){
plot.new()
if(hasArg(type)) type<-list(...)$type
else type<-"phylogram"
if(length(type)==1) type<-rep(type,2)
if(hasArg(mar)) mar<-list(...)$mar
else mar<-c(0.1,0.1,0.1,0.1)
if(hasArg(xlim)) xlim<-list(...)$xlim
else xlim<-c(-0.5,0.5)
if(hasArg(scale.bar)) scale.bar<-list(...)$scale.bar
else scale.bar<-rep(0,2)
if(hasArg(ylim)) ylim<-list(...)$ylim
else ylim<-if(any(scale.bar>0)) c(-0.1,1) else c(0,1)
if(hasArg(link.type)) link.type<-list(...)$link.type
else link.type<-"straight"
if(hasArg(link.lwd)) link.lwd<-list(...)$link.lwd
else link.lwd<-1
if(hasArg(link.col)) link.col<-list(...)$link.col
else link.col<-"black"
if(hasArg(link.lty)) link.lty<-list(...)$link.lty
else link.lty<-"dashed"
if(hasArg(edge.col)) edge.col<-list(...)$edge.col
else edge.col<-list(
left=rep("black",nrow(x$trees[[1]]$edge)),
right=rep("black",nrow(x$trees[[2]]$edge)))
if(hasArg(frame)) frame<-list(...)$frame
else frame<-c(TRUE,TRUE)
if(length(frame)==1) frame<-rep(frame,2)
obj<-list(...)
if(is.null(obj$part)) obj$part<-0.4
par(mar=mar)
plot.window(xlim=xlim,ylim=ylim)
leftArgs<-rightArgs<-obj
leftArgs$edge.col<-edge.col$left
rightArgs$edge.col<-edge.col$right
if(!is.null(obj$fsize)){
if(length(obj$fsize)>1){
leftArgs$fsize<-obj$fsize[1]
rightArgs$fsize<-obj$fsize[2]
sb.fsize<- if(length(obj$fsize)>2) obj$fsize[3] else 1
} else sb.fsize<-1
} else sb.fsize<-1
if(!is.null(obj$ftype)){
if(length(obj$ftype)>1){
leftArgs$ftype<-obj$ftype[1]
rightArgs$ftype<-obj$ftype[2]
}
}
leftArgs$frame<-frame[1]
rightArgs$frame<-frame[2]
plotter<-if(type[1]=="cladogram") "cladogram" else "phylogram"
x1<-do.call(plotter,c(list(tree=x$trees[[1]]),leftArgs))
plotter<-if(type[2]=="cladogram") "cladogram" else "phylogram"
left<-get("last_plot.phylo",envir=.PlotPhyloEnv)
x2<-do.call(plotter,c(list(tree=x$trees[[2]],direction="leftwards"),
rightArgs))
right<-get("last_plot.phylo",envir=.PlotPhyloEnv)
if(!is.null(x$assoc)) makelinks(x,c(x1,x2),link.type,link.lwd,link.col,
link.lty)
else cat("No associations provided.\n")
assign("last_plot.cophylo",list(left=left,right=right),envir=.PlotPhyloEnv)
if(any(scale.bar>0)) add.scalebar(x,scale.bar,sb.fsize)
}
## add scale bar
## written by Liam J. Revell 2015, 2023
add.scalebar<-function(obj,scale.bar,fsize){
if(scale.bar[1]>0){
pp<-get("last_plot.cophylo",envir=.PlotPhyloEnv)[[1]]
s1<-diff(range(pp$xx))/max(nodeHeights(obj$trees[[1]]))
lines(c(-0.5,-0.5+scale.bar[1]*s1),rep(-0.05,2))
lines(rep(-0.5,2),c(-0.05,-0.06))
lines(rep(-0.5+scale.bar[1]*s1,2),c(-0.05,-0.06))
text(mean(c(-0.5,-0.5+scale.bar[1]*s1)),rep(-0.05,2),scale.bar[1],pos=1)
}
if(scale.bar[2]>0){
pp<-get("last_plot.cophylo",envir=.PlotPhyloEnv)[[2]]
s2<-diff(range(pp$xx))/max(nodeHeights(obj$trees[[2]]))
lines(c(0.5-scale.bar[2]*s2,0.5),rep(-0.05,2))
lines(rep(0.5-scale.bar[2]*s2,2),c(-0.05,-0.06))
lines(rep(0.5,2),c(-0.05,-0.06))
text(mean(c(0.5-scale.bar[2]*s2,0.5)),rep(-0.05,2),scale.bar[2],pos=1)
}
}
## print an object of class "cophylo"
## written by Liam J. Revell 2015
print.cophylo<-function(x, ...){
cat("Object of class \"cophylo\" containing:\n\n")
cat("(1) 2 (possibly rotated) phylogenetic trees in an object of class \"multiPhylo\".\n\n")
cat("(2) A table of associations between the tips of both trees.\n\n")
}
## print method for "multiCophylo" object
print.multiCophylo<-function(x, ...)
cat("Object of class \"multiCophylo\" containg",length(x),"objects of class \"cophylo\".\n\n")
## written by Liam J. Revell 2015, 2016
tipRotate<-function(tree,x,...){
if(hasArg(fn)) fn<-list(...)$fn
else fn<-function(x) x^2
if(hasArg(methods)) methods<-list(...)$methods
else methods<-"pre"
if("exhaustive"%in%methods) methods<-"exhaustive"
if(hasArg(print)) print<-list(...)$print
else print<-FALSE
if(hasArg(max.exhaustive)) max.exhaustive<-list(...)$max.exhaustive
else max.exhaustive<-20
if(hasArg(rotate.multi)) rotate.multi<-list(...)$rotate.multi
else rotate.multi<-FALSE
if(rotate.multi) rotate.multi<-!is.binary(tree)
if(hasArg(anim.cophylo)) anim.cophylo<-list(...)$anim.cophylo
else anim.cophylo<-FALSE
if(anim.cophylo){
if(hasArg(left)) left<-list(...)$left
else left<-NULL
if(hasArg(right)) right<-list(...)$right
else right<-NULL
if(hasArg(assoc)) assoc<-list(...)$assoc
else assoc<-NULL
if(is.null(left)&&is.null(right)) anim.cophylo<-FALSE
if(hasArg(only.accepted)) only.accepted<-list(...)$only.accepted
else only.accepted<-TRUE
}
tree<-reorder(tree)
nn<-1:tree$Nnode+length(tree$tip.label)
if("just.compute"%in%methods){
foo<-function(phy,x) sum(fn(x-setNames(1:length(phy$tip.label),phy$tip.label)[names(x)]))
oo<-pp<-foo(tree,x)
}
if("exhaustive"%in%methods){
if(Ntip(tree)>max.exhaustive){
cat(paste("\nmethods=\"exhaustive\" not permitted for more than",
max.exhaustive,"tips.\n",
"If you are sure you want to run an exhaustive search for a tree of this size\n",
"increasing argument max.exhaustive & re-run.\n",
"Setting methods to \"pre\".\n\n"))
methods<-"pre"
} else {
cat("Running exhaustive search. May be slow!\n")
oo<-Inf
tt<-allRotations(tree)
foo<-function(phy,x) sum(fn(x-setNames(1:length(phy$tip.label),phy$tip.label)[names(x)]))
pp<-sapply(tt,foo,x=x)
ii<-which(pp==min(pp))
ii<-if(length(ii)>1) sample(ii,1) else ii
tt<-tt[[ii]]
pp<-pp[ii]
}
if(print) message(paste("objective:",pp))
tree<-tt
}
ANIM.COPHYLO<-function(tree){
dev.hold()
if(is.null(left)) plot(cophylo(tree,right,assoc=assoc,rotate=FALSE),...)
else if(is.null(right)) plot(cophylo(left,tree,assoc=assoc,rotate=FALSE),...)
nodelabels.cophylo(node=i+Ntip(tree),pie=1,col="red",cex=0.4,
which=if(is.null(left)) "left" else "right")
dev.flush()
}
if("pre"%in%methods){
for(i in 1:tree$Nnode){
if(anim.cophylo) ANIM.COPHYLO(tree)
tt<-if(rotate.multi) rotate.multi(tree,nn[i]) else untangle(rotate(tree,nn[i]),"read.tree")
oo<-sum(fn(x-setNames(1:length(tree$tip.label),tree$tip.label)[names(x)]))
if(inherits(tt,"phylo")) pp<-sum(fn(x-setNames(1:length(tt$tip.label),tt$tip.label)[names(x)]))
if(anim.cophylo&&!only.accepted) ANIM.COPHYLO(tt)
else if(inherits(tt,"multiPhylo")){
foo<-function(phy,x) sum(fn(x-setNames(1:length(phy$tip.label),phy$tip.label)[names(x)]))
pp<-sapply(tt,foo,x=x)
if(anim.cophylo&&!only.accepted) nulo<-lapply(tt,ANIM.COPHYLO)
ii<-which(pp==min(pp))
ii<-if(length(ii)>1) sample(ii,1) else ii
tt<-tt[[ii]]
pp<-pp[ii]
}
if(oo>pp) tree<-tt
if(print) message(paste("objective:",min(oo,pp)))
}
}
if("post"%in%methods){
for(i in tree$Nnode:1){
if(anim.cophylo) ANIM.COPHYLO(tree)
tt<-if(rotate.multi) rotate.multi(tree,nn[i]) else untangle(rotate(tree,nn[i]),"read.tree")
oo<-sum(fn(x-setNames(1:length(tree$tip.label),tree$tip.label)[names(x)]))
if(inherits(tt,"phylo")) pp<-sum(fn(x-setNames(1:length(tt$tip.label),tt$tip.label)[names(x)]))
if(anim.cophylo&&!only.accepted) ANIM.COPHYLO(tt)
else if(inherits(tt,"multiPhylo")){
foo<-function(phy,x) sum(fn(x-setNames(1:length(phy$tip.label),phy$tip.label)[names(x)]))
pp<-sapply(tt,foo,x=x)
if(anim.cophylo&&!only.accepted) nulo<-lapply(tt,ANIM.COPHYLO)
ii<-which(pp==min(pp))
ii<-if(length(ii)>1) sample(ii,1) else ii
tt<-tt[[ii]]
pp<-pp[ii]
}
if(oo>pp) tree<-tt
if(print) message(paste("objective:",min(oo,pp)))
}
}
attr(tree,"minRotate")<-min(oo,pp)
if(anim.cophylo) ANIM.COPHYLO(tree)
tree
}
## multi2di for "multiPhylo" object
MULTI2DI<-function(x,...){
obj<-lapply(x,multi2di,...)
class(obj)<-"multiPhylo"
obj
}
## labeling methods for plotted "cophylo" object
## written by Liam J. Revell 2015
nodelabels.cophylo<-function(...,which=c("left","right")){
obj<-get("last_plot.cophylo",envir=.PlotPhyloEnv)
if(which[1]=="left") assign("last_plot.phylo",obj[[1]],envir=.PlotPhyloEnv)
else if(which[1]=="right") assign("last_plot.phylo",obj[[2]],envir=.PlotPhyloEnv)
nodelabels(...)
}
edgelabels.cophylo<-function(...,which=c("left","right")){
obj<-get("last_plot.cophylo",envir=.PlotPhyloEnv)
if(which[1]=="left") assign("last_plot.phylo",obj[[1]],envir=.PlotPhyloEnv)
else if(which[1]=="right") assign("last_plot.phylo",obj[[2]],envir=.PlotPhyloEnv)
edgelabels(...)
}
tiplabels.cophylo<-function(...,which=c("left","right")){
obj<-get("last_plot.cophylo",envir=.PlotPhyloEnv)
if(which[1]=="left") assign("last_plot.phylo",obj[[1]],envir=.PlotPhyloEnv)
else if(which[1]=="right") assign("last_plot.phylo",obj[[2]],envir=.PlotPhyloEnv)
tiplabels(...)
}
## function to draw sigmoidal links
## modified from https://stackoverflow.com/questions/32046889/connecting-two-points-with-curved-lines-s-ish-curve-in-r
drawCurve<-function(x,y,scale=0.01,...){
x1<-x[1]
x2<-x[2]
y1<-y[1]
y2<-y[2]
curve(plogis(x,scale=scale,location=(x1+x2)/2)*(y2-y1)+y1,
x1,x2,add=TRUE,...)
}
## S3 summary method
## written by Liam J. Revell 2016
summary.cophylo<-function(object,...){
cat("\nCo-phylogenetic (\"cophylo\") object:",deparse(substitute(object)),
"\n\n")
cat(paste("Tree 1 (left tree) is an object of class \"phylo\" containing",
Ntip(object$trees[[1]]),"species.\n\n"))
cat(paste("Tree 2 (right tree) is an object of class \"phylo\" containing",
Ntip(object$trees[[2]]),"species.\n\n"))
cat("Association (assoc) table as follows:\n\n")
maxl<-max(sapply(strsplit(object$assoc[,1],""),length))
cat(paste("\tleft:",paste(rep(" ",max(0,maxl-5)),collapse=""),
"\t----\tright:\n",sep=""))
nulo<-apply(object$assoc,1,function(x,maxl) cat(paste("\t",x[1],
paste(rep(" ",maxl-length(strsplit(x[1],split="")[[1]])),
collapse=""),"\t----\t",x[2],"\n",sep="")),maxl=maxl)
cat("\n")
}
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